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23 May 2022

Development and Assessment of Equine Behaviour


This chapter describes the process of evaluating equine behaviour during domestication, learning, in cases with clinical conditions, and on equine welfare.

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Appleby M.C. and Hughes B.O. (eds) (1997) Animal Welfare . CAB International, Wallingford, UK.
Back W. and Clayton H.M. (2013)  Equine Locomotion-E-Book . Elsevier Health Sciences.
Bateson P. (2003) The promise of behavioural biology. Animal Behaviour 65, 11–17.
Bateson P. and Horn G. (1994) Imprinting and recognition memory: a neural net model. Animal Behaviour 48, 695–715.
Budiansky S. (1997) The Nature of Horses . The Free Press, New York, pp. 211–234.
Christiansen S.B. and Forkman B. (2007) Assessment of animal welfare in a veterinary context – a call for ethologists. Applied Animal Behaviour Science 106, 203–220.
Clutton-Brock J. (1992) Horse Power: A History of the Horse and the Donkey in Human Societies . Harvard University Press, Cambridge, MA, pp. 192.
Clutton-Brock J. (1999) A Natural History of Domesticated Mammals . Cambridge University Press, Cambridge.
Dalla Costa E., Minero M., Lebelt D., Stucke D., Canali E. and Leach M.C. (2014) Development of the horse grimace scale (HGS) as a pain assessment tool in horses undergoing routine castration. PLoS ONE 9(3), e92281.
Dawkins M.S. (2004) Using behaviour to assess welfare. Animal Welfare 13, 53–57.
Domjam M. (1998) The Principles of Learning and Behaviour , 4th edn. Brooks/Cole, London.
Forkman B.A. (2002) Learning and cognition. In: Jensen P. (ed.) The Ethology of Domestic Animals . CAB International, Wallingford, UK, pp. 51–64.
Fraser A.F. (2008) Veterinarians and animal welfare – a comment. The Canadian Veterinary Journal 49(1), 8.
Goodwin D., McGreevy P., Waran N. and McLean A. (2009) How equitation science can elucidate and refine horsemanship techniques. Veterinary Journal 181, 5–11.
Gore T., Gore P. and Griffin J.M.L. (2008) Horse Owner’s Veterinary Handbook , 3rd edn. Wiley Publishing Inc., Hoboken, New Jersey.
Hanggi E.B. ( 2003). Discrimination learning based on relative size concepts in horses (Equus caballus). Applied Animal Behaviour Science 83(3), 201– 213.
Heleski C., Bauson L. and Bello N. (2008) Evaluating the addition of positive reinforcement for learning a frightening task: a pilot study with horses. Journal of Applied Animal Welfare Science 11(3), 213–222.
Henry S., Fureix C., Rowberry R., Bateson M. and Hausberger M. (2017) Do horses with poor welfare show ‘pessimistic’ cognitive biases?  Naturwissenschaften  104, 8.
Herdt T.H. (2007) Gastrointestinal physiology and metabolism (Section IV). In: Cunningham J.G. and Klein B.G. (eds) Veterinary Physiology . Saunders Elsevier, Amsterdam.
Houpt K.A. (1991) Domestic Animal Behaviour for Veterinarians and Animal Scientists , 2nd edn. Iowa State University Press, Ames, Iowa.
Jensen P. (2001) Parental behaviour. In: Keeling L.J. and Gonyou H.W. (eds) Social Behaviour in Farm Animals . CAB International, Wallingford, UK, pp. 59–82.
Jones S. (2001) Darwin’s Ghost . The Origin of Species Updated. Anchor Canada, Random House of Canada Ltd, Toronto.
Jørgensen G.H., Aanensen L., Mejdell C.M. and Bøe K.E. (2016) Preference for shelter and additional heat in horses exposed to Nordic winter conditions. Equine Veterinary Journal 48, 720–726.
Kiley-Worthington M. (1999) The Behaviour of Horses . J.A. Allen & Co. Ltd, London.
Kilgour R. and Dalton C. (1984) Livestock Behaviour: A Practical Guide , 1st edn. CRC Press, New York, pp. 334.
Lansade L., Bouissue M.F. and Erhard H.W. (2008) Fearfulness in horses: a temperamental trait, stable across time and situations. Applied Animal Behaviour Science 115, 182–200.
Lehner P.N. (1996) Handbook of Ethological Methods , 2nd edn. Cambridge University Press, Cambridge.
Ligout S., Bouissou M.F. and Boiviu X. (2008) Comparison of the effects of two different handling methods on the subsequent behaviour of Anglo-Arabian foals towards humans and handling. Applied Animal Behaviour Science 113, 175–188.
Lloyd A.S., Martin J.E., Bornett-Gauci H.L.I. and Wilkinson R.G. (2008) Horse personality: variation between breeds. Applied Animal Behaviour Science 112, 369–383.
Lorenz K. (1965) Evolution and Modification of Behaviour . University of Chicago Press, Chicago, Illinois.
Martin P. and Bateson P. (2007) Measuring Behaviour , 2nd edn. Cambridge University Press, Cambridge.
McDonnell S. (2003) A Practical Field Guide to Horse Behavior: The Equid Ethogram . The Blood Horse Inc., Lexington, KY.
McGreevy P. (2004) Equine Behaviour . Saunders, Edinburgh, UK.
Miller R. (2003) Imprint Training of the Newborn Foal . Western Horseman Publishing.
Mills N.J. (1991) Mound making in azoturia cases. Veterinary Record 128, 215.
Morresey P.R. (2005) Prenatal and perinatal indicators of neonatal viability. Clinical Techniques in Equine Practice 4, 238–249.
Oliver J.E. and Lorenz M.D. (1993) Handbook of Veterinary Neurology , 2nd edn. W.B. Saunders, Philadelphia, Pennsylvania.
Pearson G. (2016) Equine learning theory and the horse-veterinarian interaction. MScR thesis. University of Edinburgh, UK.
Riolo R.L., Cohen M.D. and Axeford R. (2001) Evolution of cooperation without reciprocity. Nature 414, 441–443.
Robinson N.E. (2007) Respiratory function (Section VIII) and homeostasis (Section IX). In: Cunningham J.G. and Klein B.G. (eds) Veterinary Physiology . Saunders Elsevier, Amsterdam.
Russell C.M. and Wilkins P.A. (2006) Evaluation of the recumbent neonate . Clinical Techniques in Equine Practice 5, 161–171.
Sambraus H.H. (1985) Mouth-based anomalous syndromes. In: Fraser A.F. (ed.) Ethology of Farm Animals, World Animal Science . Elsevier, Amsterdam, pp. 381–411.
Sankey C., Richard-Yris M.A., Henry S., Fureix C., Nassur F. and Hausberger M. (2010) Reinforcement as a mediator of the perception of humans by horses. Animal Cognition 13, 753–765.
Seaman S.C., Davidson H.P.B. and Waran N.K. (2002) How reliable is temperament assessment in the domestic horse (Equus caballus)? Applied Animal Behaviour Science 78, 157–191.
Stubbs N., Menke, E. Back, W. and Clayton H.M. (2013) Rehabilitation of locomotor apparatus. In: Back W. and Clayton H.M. (eds) Equine Locomotion , 2nd edn. Saunders Elsevier, Edinburgh, UK, pp. 381–418.
Visser E.K., van Reenen C.G., Hopster H., Schilder M.B.H., Knaap J.H. et al. (2001) Quantifying aspects of young horses’ temperament. Consistency of behavioural variables. Applied Animal Behaviour Science 74, 241–258.
Visser E.K., Van Reenen C.G., Schilder M.B.H., Barneveld A. and Blokhuis H.J. (2003) Learning performances in young horses using two different learning tests. Applied Animal Behaviour Science 80, 311–326.
Waran N. and Randle H. (2017) What we can measure, we can manage: The importance of using robust welfare indicators in Equitation Science. Applied Animal Behaviour Science 190, 74–81.
Waran N.K., Clarke N. and Farnworth M. (2008) The effects of weaning on the domestic horse (Equus caballus) Applied Animal Behaviour Science 110(1–2), 42–57.
Willeberg P. (1997) Epidemiology and animal welfare. Epidemiologie. Santé Animale 31, 3–7.
Zentall T.R. (1996) The analysis of initiative learning in animals. In: Hayes C.M. and Galef B.G. (eds) Social Learning of Animals: The Roots of Culture . Academic Press, San Diego, California, pp. 221–243.

Information & Authors


Published In

Pages: 1 - 23
Editors: Christopher B Riley, Massey University, New Zealand, Sharon E Cregier, University of Prince Edward Island, Canada and North American Editor Emerita, Equine Behaviour, and Andrew Fraser, formerly Memorial University of Newfoundland, Canada
ISBN (ePDF): 978-1-78924-212-6
ISBN (ePUB): 978-1-78924-213-3
ISBN (hardback): 978-1-78924-210-2
ISBN (paperback): 978-1-78924-211-9


Cover date: 17 May 2022
Published online: 23 May 2022





Natalie K. Waran, PhD [email protected]
Eastern Institute of Technology, Hawke’s Bay, New Zealand;
Gemma Pearson, BVMS, MScR, MRCVS
The University of Edinburgh, Midlothian, Scotland, UK
Andrew F. Fraser, MVSc, MRCVS
Carbonear, Newfoundland, Canada


*Corresponding author: [email protected]

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