Fallopia convolvulus (black bindweed)
Datasheet Types: Pest, Invasive species, Host plant
Abstract
This datasheet on Fallopia convolvulus covers Identity, Overview, Distribution, Dispersal, Hosts/Species Affected, Diagnosis, Biology & Ecology, Environmental Requirements, Natural Enemies, Impacts, Uses, Prevention/Control, Further Information.
Identity
- Preferred Scientific Name
- Fallopia convolvulus (L.) Á. Löve
- Preferred Common Name
- black bindweed
- Other Scientific Names
- Bilderdykia convolvulus (L.) Dumort.
- Fagopyrum convolvulus (L.) H. Gross
- Fagopyrum volubile Gilib.
- Fallopia convolvulus var. subalatum (Lej. & Court.) D.H.Kent
- Helxine convolvulus (L.) Raf.
- Polygonum convolvulus L.
- Reynoutria convolvulus (L.) Shinners
- Tiniaria convolvulus L. Webb. & Moq.
- International Common Names
- Englishbear-bindbind-cornclimbing bindweedclimbing buckwheatwild buckwheat
- Spanishchilillocorregüela anualpoligono trepador
- Frenchfaux liseronrenouée liseronvrillée sauvage
- Portuguesecipo de veado de invernocorriola-bastarda
- Local Common Names
- Argentinaenredadera
- Brazilcipo-de-veado-de-inverno
- Chileenredadera
- Denmarksnerle-pileurt
- Finlandkiertotatar
- GermanyGemeiner Winden-KnöterichWinden-Knöterich
- Iranpichak band
- Italyconvolvolo neroerba leprinapoligono convolvolo
- Japansobakazura
- Moroccofaux liseron
- Netherlandswilde boekweitzwaluwtong
- New Zealandcornbind
- Norwayvindelskjedekne
- Swedenaakerbinda
- Turkeysarmasik coban
- EPPO code
- POLCO (Polygonum convolvulus)
Pictures

Habit
Fallopia convolvulus (black bindweed); habit, in a rock crevice. Bigfork, Montana, USA. July 2012.
©Prof Matt Lavin-2009/Bozeman, Montana, USA - CC BY-SA 2.0

Habit
Fallopia convolvulus (black bindweed); habit, in a rock crevice. Bigfork, Montana, USA. July 2012.
©Prof Matt Lavin-2009/Bozeman, Montana, USA - CC BY-SA 2.0

Leaf
Fallopia convolvulus (black bindweed); leaf. Bigfork, Montana, USA. July 2012.
©Prof Matt Lavin-2009/Bozeman, Montana, USA - CC BY-SA 2.0

Leaf and flower
Fallopia convolvulus (black bindweed); leaf and flower. Bigfork, Montana, USA. July 2012.
©Prof Matt Lavin-2009/Bozeman, Montana, USA - CC BY-SA 2.0

Leaf and flower
Fallopia convolvulus (black bindweed); leaf and flower. Bigfork, Montana, USA. July 2012.
©Prof Matt Lavin-2009/Bozeman, Montana, USA - CC BY-SA 2.0

Flower
Fallopia convolvulus (black bindweed); close-up of flower. Bigfork, Montana, USA. July 2012.
©Prof Matt Lavin-2009/Bozeman, Montana, USA - CC BY-SA 2.0

Habit
Fallopia convolvulus (black bindweed); habit showing flowering plant. Leaves are 2-6cm long, with small and inconspicuous flowers to 5mm in diameter.
©Chris Parker/Bristol, UK
Summary of Invasiveness
F. convolvulus is a weedy species of gardens, cultivated fields, open habitats, orchards, non-crop areas, waste areas, and disturbed sites. It is well-adapted to a wide range of climatic conditions and soils. This species is a prolific seed producer and has the potential to produce up to 30,000 seeds/plant. Seeds can be dispersed by farm machinery, and water. It is also a common contaminant of wheat and other cereal crops.
F. convolvulus is often a serious weed in cereals, vegetables and horticultural crops (FAO, 2015). Currently, it is listed as invasive in the Dominican Republic, Cuba, Australia, New Caledonia, and New Zealand (Webb et al., 1988; MacKee, 1994; Wilson, 2008; Acevedo-Rodriguez and Strong, 2012), but it is also ranked as a serious weed in 20 crops in more than 41 countries around the world (Holm et al., 1991).
F. convolvulus is often a serious weed in cereals, vegetables and horticultural crops (FAO, 2015). Currently, it is listed as invasive in the Dominican Republic, Cuba, Australia, New Caledonia, and New Zealand (Webb et al., 1988; MacKee, 1994; Wilson, 2008; Acevedo-Rodriguez and Strong, 2012), but it is also ranked as a serious weed in 20 crops in more than 41 countries around the world (Holm et al., 1991).
Taxonomic Tree
Notes on Taxonomy and Nomenclature
Polygonaceae is a well-defined family of flowering plants including around 59 genera and 1580 species (the Plant List, 2013). Members of the Polygonaceae are very diverse in habit ranging from annual and perennial herbs to shrubs, lianas and trees. The most distinctive feature of the family is the presence of an “ocrea”, a membranous or hyaline sheath uniting the stipules (Maharajan and Rajendran, 2014). The genus Polygonum sensu lato was circumscribed by Linnaeus very broadly, causing further authors to divide it into several more naturally circumscribed genera (Holub, 1970). The genus Fallopia is often included in a broader concept of Polygonum but is distinguished by a syndrome of anatomical and morphological characters (Decraene et al., 2000). Molecular data confirm its close relationship to Polygonum in the narrow sense (Lamb Frye and Kron, 2003; Freeman and Hinds, 2005).
Plant Type
Annual
Herbaceous
Perennial
Seed propagated
Description
F. convolvulus is an annual or perennial climbing herb with a thin, spindle-formed and deep root, which is often profusely branched. The stem is slender, 5-250 cm long, with long internodes. It is freely branched from the base, smooth to slightly rough, greenish, sometimes with a reddish tinge, trailing on the ground or twining around other plants. The leaves are alternate, 2-6 cm long, long-petioled, elongate-ovate, pointed, heart- or arrow-shaped. The stipule sheath or ochrea with smooth margins. Flowers are small, inconspicuous, up to 5 mm in diameter, and grouped in short axillary clusters of 2 to 6 flowers or in terminal interrupted or spike-like racemes. The perianth is reddish green, white inside and along the margins; the short pedicels are articulate near the upper end. The fruit is a triangular achene, 3-4 mm long, with an obtuse base and pointed top, minutely pitted, brownish black, dull, after maturity enclosed by the somewhat enlarged outer perigon leaves (Korsmo, 1954; Holm et al., 1991; Conert et al., 1981).
The seedlings have long cotyledons, 7-33 mm in length. They are four times longer than wide, with obtuse points. The upper leaf surface is dull dark green; the lower leaf surface is light green, with a distinct central nerve. The expanded cotyledons generally assume a 120° angle, rather than being opposite, at the point of which the primary leaf appears. At the beginning, the primary leaf is often laterally rolled up, with a blueish or reddish green tinge (Schwär et al., 1970; Hume et al., 1983).
The seedlings have long cotyledons, 7-33 mm in length. They are four times longer than wide, with obtuse points. The upper leaf surface is dull dark green; the lower leaf surface is light green, with a distinct central nerve. The expanded cotyledons generally assume a 120° angle, rather than being opposite, at the point of which the primary leaf appears. At the beginning, the primary leaf is often laterally rolled up, with a blueish or reddish green tinge (Schwär et al., 1970; Hume et al., 1983).
Distribution
F. convolvulus is native to Eurasia, although due to its plasticity it occurs in most regions of the world. In Europe and North America it grows everywhere that crops are cultivated, but is less frequent in the south (Franzini, 1982). In Canada, it is reported as one of the most abundant weeds, occurring in 60-80% of all fields in the provinces of Alberta, Manitoba, Saskatchewan and Prince Edward Island (Hume et al., 1983). It is found throughout South America. In Africa it occurs in North Africa, on the east coast and in South Africa. In Asia, its geographical distribution ranges from Japan to Iran, down to India and into Indonesia. F. convolvulus can also be found in Australia and New Zealand. It is not common in the humid tropics, and in warm areas, it is more often found at high altitudes or in cooler valleys (Holm et al., 1991).
Distribution Map
Distribution Table
Means of Movement and Dispersal
F. convolvulus spreads by seeds and may produce 11,900-30,000 seeds/plant (Stevens, 1932; Forsberg and Best, 1964). The hard seed coat allows seeds to remain dormant for several years (Chippendale and Milton, 1934, Roberts and Feast, 1973, Conn and Deck, 1995). Seeds can be dispersed by farm machinery and water over short distances. It is also a common contaminant of wheat and other cereal crops (Gooch, 1963, Rutledge and McLendon, 1996).
Pathway Causes
Pathway cause | Notes | Long distance | Local | References |
---|---|---|---|---|
Crop production (pathway cause) | Contaminant in cereals crops | Yes | Yes |
Pathway Vectors
Pathway vector | Notes | Long distance | Local | References |
---|---|---|---|---|
Debris and waste associated with human activities (pathway vector) | Yes | Yes | ||
Water (pathway vector) | Seed | Yes | ||
Wind (pathway vector) | Yes |
Hosts/Species Affected
A list of crops in which F. convolvulus is, or could be, a problem weed includes almost every crop of the temperate zone. Worldwide, F. convolvulus is most troublesome in cereals, but it may also cause yield losses in potatoes, sugarbeet and vegetables, as well as vineyards and orchards. According to Holm et al. (1991), it is a weed of 25 crops in 41 countries and in 20 crops of these countries it is ranked as a serious weed.
Host Plants and Other Plants Affected
Host | Family | Host status | References |
---|---|---|---|
Allium cepa (onion) | Liliaceae | Other | |
Asparagus officinalis (asparagus) | Liliaceae | Other | |
Avena sativa (oats) | Poaceae | Main | |
Beta vulgaris var. saccharifera (sugarbeet) | Chenopodiaceae | Main | |
Brassica | Brassicaceae | Main | |
Brassica napus var. oleifera | Brassicaceae | Unknown | |
Daucus carota (carrot) | Apiaceae | Other | |
Glycine max (soyabean) | Fabaceae | Main | |
Gossypium (cotton) | Malvaceae | Other | |
Helianthus annuus (sunflower) | Asteraceae | Other | |
Hordeum vulgare (barley) | Poaceae | Main | |
Linum usitatissimum (flax) | Main | ||
Medicago (medic) | Fabaceae | Other | |
Medicago sativa (lucerne) | Fabaceae | Unknown | |
Phaseolus vulgaris (common bean) | Fabaceae | Other | |
Pistacia vera (pistachio) | Anacardiaceae | Unknown | |
Pisum sativum (pea) | Fabaceae | Other | |
Secale cereale (rye) | Poaceae | Main | |
Solanum lycopersicum (tomato) | Solanaceae | Unknown | |
Solanum tuberosum (potato) | Solanaceae | Main | |
Spinacia oleracea (spinach) | Chenopodiaceae | Other | |
Triticum aestivum (wheat) | Poaceae | Main | |
Triticum turgidum subsp. durum | Poaceae | Unknown | |
Zea mays (maize) | Poaceae | Main |
Similarities to Other Species/Conditions
Prior to flowering, F. convolvulus may be mistaken for Convolvulus arvensis, as both have twining greenish stems and leaves of similar shape. In contrast to F. convolvulus, C. arvensis is a deep-rooted perennial with extensive creeping white roots and rhizomes, blunt-tipped leaves and large funnel-shaped pink or white flowers. It has no ochrea or sheathing stipules as in F. convolvulus.
Two further Fallopia species which could be confused with F. convolvulus in Canada (Hume et al. (1983): Fallopia cilinode and F. scandens. Both are perennials. Fallopia cilinode has bristles at the base of the sheath, leaves with narrower spacing between the basal lobes and achenes that are shiny and smooth. The flowers of F. scandens are long-stalked. It has a strong, winged calyx and smooth, shiny achenes.
F. convolvulus is also closely related to Fallopia dumetorum and F. dentate-alata, but in both these species the pedicel is much longer (up to 10 mm long) and articulated above the middle (Flora of Pakistan Editorial Committee, 2013).
Two further Fallopia species which could be confused with F. convolvulus in Canada (Hume et al. (1983): Fallopia cilinode and F. scandens. Both are perennials. Fallopia cilinode has bristles at the base of the sheath, leaves with narrower spacing between the basal lobes and achenes that are shiny and smooth. The flowers of F. scandens are long-stalked. It has a strong, winged calyx and smooth, shiny achenes.
F. convolvulus is also closely related to Fallopia dumetorum and F. dentate-alata, but in both these species the pedicel is much longer (up to 10 mm long) and articulated above the middle (Flora of Pakistan Editorial Committee, 2013).
Habitat
F. convolvulus is most common on arable land, but it can also be found on waste ground, in thickets, on roadsides, along fences, and occasionally in pastures and on river banks (Hume et al., 1993). Because of its deep root system, it is rather unaffected by drought or low nitrogen levels. Nevertheless, high nutrient levels promote the growth and abundance of F. convolvulus (Bazdyrev et al., 1984; Mahn, 1984; Pulcher-Häussling and Hurle, 1986; Borowiec and Kutyna, 1988; Bräutigam, 1995). According to Karczmarczyk et al. (1983), who investigated the influence of irrigation on potato, sugar-beet and the weeds associated with these crops, F. convolvulus predominates on non-irrigated field plots. Haas and Streibig (1982) and Haman and Peeper (1983) reported that growth of F. convolvulus is stimulated by shading.
While in northern areas it is found in warm and well-drained locations, in hotter countries it is present on moist shady sites and at high altitudes. In China, F. convolvulus is found in thickets in valleys and along stream banks, at 100-3600 m (Flora of China Editorial Committee, 2015). In Pakistan, as well as being a weed on cultivated land, it is recorded in crevices in moist, shady places and at 1500-3500 m altitude (Flora of Pakistan Editorial Committee, 2013).
While in northern areas it is found in warm and well-drained locations, in hotter countries it is present on moist shady sites and at high altitudes. In China, F. convolvulus is found in thickets in valleys and along stream banks, at 100-3600 m (Flora of China Editorial Committee, 2015). In Pakistan, as well as being a weed on cultivated land, it is recorded in crevices in moist, shady places and at 1500-3500 m altitude (Flora of Pakistan Editorial Committee, 2013).
Habitat List
Category | Sub category | Habitat | Presence | Status |
---|---|---|---|---|
Terrestrial | ||||
Terrestrial | Terrestrial – Managed | Cultivated / agricultural land | Present, no further details | Harmful (pest or invasive) |
Terrestrial | Terrestrial – Managed | Cultivated / agricultural land | Present, no further details | Natural |
Terrestrial | Terrestrial – Managed | Managed forests, plantations and orchards | Present, no further details | Harmful (pest or invasive) |
Terrestrial | Terrestrial – Managed | Managed forests, plantations and orchards | Present, no further details | Natural |
Terrestrial | Terrestrial – Managed | Managed grasslands (grazing systems) | Present, no further details | Harmful (pest or invasive) |
Terrestrial | Terrestrial – Managed | Managed grasslands (grazing systems) | Present, no further details | Natural |
Terrestrial | Terrestrial – Managed | Disturbed areas | Present, no further details | Harmful (pest or invasive) |
Terrestrial | Terrestrial – Managed | Disturbed areas | Present, no further details | Natural |
Terrestrial | Terrestrial – Managed | Rail / roadsides | Present, no further details | Harmful (pest or invasive) |
Terrestrial | Terrestrial – Managed | Rail / roadsides | Present, no further details | Natural |
Terrestrial | Terrestrial ‑ Natural / Semi-natural | Natural grasslands | Present, no further details | Harmful (pest or invasive) |
Terrestrial | Terrestrial ‑ Natural / Semi-natural | Natural grasslands | Present, no further details | Natural |
Biology and Ecology
Genetics
F. convolvulus is polyploid and has a chromosome count of 2n = 40 (Mulligan, 1957, 1960; Clapham et al., 1962).
F. convolvulus is polyploid and has a chromosome count of 2n = 40 (Mulligan, 1957, 1960; Clapham et al., 1962).
Reproductive Biology
Flowers of F. convolvulus are bisexual and self-compatible (Mulligan and Findlay, 1970). Each flower produces a single achene. In Asia, this species has been recorded flowering from July to September (FAO, 2015). In Europe, flowering plants can be found throughout the summer till mid-autumn (Hanf, 1982). As a result of the indeterminate flowering habit of F. convolvulus, flowers, immature seed and mature seed may be found at the same time on a single plant.
Flowers of F. convolvulus are bisexual and self-compatible (Mulligan and Findlay, 1970). Each flower produces a single achene. In Asia, this species has been recorded flowering from July to September (FAO, 2015). In Europe, flowering plants can be found throughout the summer till mid-autumn (Hanf, 1982). As a result of the indeterminate flowering habit of F. convolvulus, flowers, immature seed and mature seed may be found at the same time on a single plant.
Longevity
F. convolvulus is a fast-growing herb that behaves as annual and perennial herb. It has a large, fibrous root system which may reach down as far as 80 cm in the soil (Kutschera, 1960), enabling it to evade dry conditions.
F. convolvulus is a fast-growing herb that behaves as annual and perennial herb. It has a large, fibrous root system which may reach down as far as 80 cm in the soil (Kutschera, 1960), enabling it to evade dry conditions.
Physiology and Phenology
According to Hanf (1982), a single plant of F. convolvulus produces 100 to 1000 seeds. Grown without competition, Stevens (1932) reported fecundity of 11,900 seeds per plant, and Forsberg and Best (1964) obtained up to 30,000 seeds from plants that emerged early in the growing season. The seeds possess a deep primary dormancy for several months when mature (Zwerger, 1987, 1993) due mostly to the hard pericarp, which limits gas and water exchange, and acts as a barrier to germination (Ransom, 1935; Timson, 1966). Under field conditions, most seeds germinate in their first year (Chepil, 1946), but they may remain viable in the soil for several years (Holm et al., 1991). Chippendale and Milton (1934) found viable seeds of F. convolvulus after approximately 22 years in the soil under pasture.
Seedlings emerge throughout the growing season. They normally germinate at depths in the soil between 6 and 51 mm, although research has shown that seeds buried as deep as 19 cm can germinate (Forsberg and Best 1964). Light is not required for germination. Seeds germinate at temperatures between 2°C and 30°C, with maximum germination rates occurring between 5°C and 15°C.
According to Hanf (1982), a single plant of F. convolvulus produces 100 to 1000 seeds. Grown without competition, Stevens (1932) reported fecundity of 11,900 seeds per plant, and Forsberg and Best (1964) obtained up to 30,000 seeds from plants that emerged early in the growing season. The seeds possess a deep primary dormancy for several months when mature (Zwerger, 1987, 1993) due mostly to the hard pericarp, which limits gas and water exchange, and acts as a barrier to germination (Ransom, 1935; Timson, 1966). Under field conditions, most seeds germinate in their first year (Chepil, 1946), but they may remain viable in the soil for several years (Holm et al., 1991). Chippendale and Milton (1934) found viable seeds of F. convolvulus after approximately 22 years in the soil under pasture.
Seedlings emerge throughout the growing season. They normally germinate at depths in the soil between 6 and 51 mm, although research has shown that seeds buried as deep as 19 cm can germinate (Forsberg and Best 1964). Light is not required for germination. Seeds germinate at temperatures between 2°C and 30°C, with maximum germination rates occurring between 5°C and 15°C.
Environmental Requirements
F. convolvulus occurs on a wide range of soil types (Hume et al., 1983), but according to Paatela and Erviö (1971) it is less common on peaty soils. In field observations in Finland, F. convolvulus was more abundant in clay than in coarse mineral or organic soil (Erviö et al., 1994). Higher rates of F. convolvulus emergence are recorded on soils with a high clay content. In field experiments in Poland, F. convolvulus predominated on plots fertilized with nitrogen plus calcium (Borowiec et al., 1995). However, Zwerger (1990) could only find a positive effect of nitrogen fertilization on seed production at low densities of this weed.
At higher weed densities, the effect of intraspecific competition became more important than nitrogen supply. Sirbu and Slonovschi (1989) found that nutrient reserves of F. convolvulus seeds increased with increasing phosphate fertilization. In a study concerning the distribution of several common weeds in Denmark, decreasing potassium content in the soil appeared to favour the occurrence of F. convolvulus (Andreasen and Streibig, 1990; Andreasen et al., 1991). According to Hanf (1982), it is one of the most frequent weeds on soils with a low pH.
Shade usually suppresses the growth of black bindweed (Haman and Peeper 1983), but in hotter countries it can be found in moist shady places and crevices (Flora of Pakistan Editorial Committee, 2013).
F. convolvulus occurs on a wide range of soil types (Hume et al., 1983), but according to Paatela and Erviö (1971) it is less common on peaty soils. In field observations in Finland, F. convolvulus was more abundant in clay than in coarse mineral or organic soil (Erviö et al., 1994). Higher rates of F. convolvulus emergence are recorded on soils with a high clay content. In field experiments in Poland, F. convolvulus predominated on plots fertilized with nitrogen plus calcium (Borowiec et al., 1995). However, Zwerger (1990) could only find a positive effect of nitrogen fertilization on seed production at low densities of this weed.
At higher weed densities, the effect of intraspecific competition became more important than nitrogen supply. Sirbu and Slonovschi (1989) found that nutrient reserves of F. convolvulus seeds increased with increasing phosphate fertilization. In a study concerning the distribution of several common weeds in Denmark, decreasing potassium content in the soil appeared to favour the occurrence of F. convolvulus (Andreasen and Streibig, 1990; Andreasen et al., 1991). According to Hanf (1982), it is one of the most frequent weeds on soils with a low pH.
Shade usually suppresses the growth of black bindweed (Haman and Peeper 1983), but in hotter countries it can be found in moist shady places and crevices (Flora of Pakistan Editorial Committee, 2013).
Climate
Climate type | Description | Preferred or tolerated | Remarks |
---|---|---|---|
Af - Tropical rainforest climate | > 60mm precipitation per month | Tolerated | |
Am - Tropical monsoon climate | Tropical monsoon climate ( < 60mm precipitation driest month but > (100 - [total annual precipitation(mm}/25])) | Tolerated | |
As - Tropical savanna climate with dry summer | < 60mm precipitation driest month (in summer) and < (100 - [total annual precipitation{mm}/25]) | Tolerated | |
Aw - Tropical wet and dry savanna climate | < 60mm precipitation driest month (in winter) and < (100 - [total annual precipitation{mm}/25]) | Tolerated | |
Cf - Warm temperate climate, wet all year | Warm average temp. > 10°C, Cold average temp. > 0°C, wet all year | Tolerated | |
Cs - Warm temperate climate with dry summer | Warm average temp. > 10°C, Cold average temp. > 0°C, dry summers | Tolerated | |
Cw - Warm temperate climate with dry winter | Warm temperate climate with dry winter (Warm average temp. > 10°C, Cold average temp. > 0°C, dry winters) | Tolerated | |
Df - Continental climate, wet all year | Continental climate, wet all year (Warm average temp. > 10°C, coldest month < 0°C, wet all year) | Preferred | |
Ds - Continental climate with dry summer | Continental climate with dry summer (Warm average temp. > 10°C, coldest month < 0°C, dry summers) | Preferred | |
Dw - Continental climate with dry winter | Continental climate with dry winter (Warm average temp. > 10°C, coldest month < 0°C, dry winters) | Preferred |
Soil Tolerances
Soil texture > light
Soil texture > medium
Soil reaction > acid
List of Pests
Notes on Natural Enemies
Insects that have been cited as using F. convolvulus as a host are Gastrophysa polygoni (Forsberg, 1955; Kjaer and Elmegaard, 1996), Coleophora therinella and C. peribenanderi (van der Wolf, 1992). F. convolvulus may serve as a habitat for a number of viruses, insects (Mamestra (Noctuidae) and Pegomyia (Anthomyiidae)), nematodes (species of Heterodera and Meloidogyne) and fungi (species of Ustilago, Puccinia and Peronospora). A review of arthropod natural enemies in Canada and Europe was presented by Hume (1983).
In a study on the internal and surface-dwelling pathogenic fungi of seeds of the 30 most widespread weed species in Lithuania, Grigaliunaite and Kacergius (1995) found that F. convolvulus seeds were the least damaged of all weeds investigated.
In a study on the internal and surface-dwelling pathogenic fungi of seeds of the 30 most widespread weed species in Lithuania, Grigaliunaite and Kacergius (1995) found that F. convolvulus seeds were the least damaged of all weeds investigated.
Natural enemies
Natural enemy | Type | Life stages | Specificity | References | Biological control in | Biological control on |
---|---|---|---|---|---|---|
Coleophora peribenanderi | Herbivore | Leaves | ||||
Coleophora therinella | Herbivore | Leaves | ||||
Gastrophysa polygoni (leaf beetle, knotweed) | Herbivore | Leaves | ||||
Pegomya setaria | Herbivore | Leaves | ||||
Peronospora polygoni | Pathogen | Leaves | ||||
Puccinia polygoni-amphibii | Pathogen | Leaves |
Impact Summary
Category | Impact |
---|---|
Economic/livelihood | Positive and negative |
Environment (generally) | Positive and negative |
Impact
F. convolvulus can reduce crop yields by competition, especially in highly infested fields (Friesen and Shebeski, 1960; Nakoneshny and Friesen, 1961; Dosland and Arnold, 1966; Fabricius and Nalewaja, 1968). Friesen and Shebeski (1960) showed that 56 and 210 plants per m² can reduce wheat yields by 15 and 25%, respectively. Crop seed weight and protein contents can also be negatively affected (Nakoneshny and Friesen, 1961; Gruenhagen and Nalewaja, 1969).
By climbing up the crop, F. convolvulus causes lodging in grain crops (Neururer, 1961; Hume et al., 1983), and can cause harvesting problems when its vines wrap around moving parts of machinery (Forsberg and Best, 1964; Fabricius and Nalewaja, 1968).
In addition, high weed densities can raise the moisture content of harvested grain (Neururer, 1961) and contribute to heating in storage when harvested with cereals as a seed contaminant (Holm et al., 1991). F. convolvulus produces large amounts of seed and they are often difficult to separate from grain crops, because of their similar size. Therefore, F. convolvulus is a serious contaminant of seed stocks in several places in the world (Gooch, 1963; Bogdan, 1965).
The general importance of this weed is due to its ability to emerge throughout the growing season. Forsberg and Best (1964) showed that late-emerging seedlings are likely to escape herbicide spraying and could be a potential source of re-infestations, especially where competition from other weeds has been eliminated.
Furthermore, F. convolvulus may serve as an alternate host for disease organisms affecting crops (Cooper and Harrison, 1973; Bendixen et al., 1979).
According to Holm et al. (1991), F. convolvulus is ranked as a principal or serious weed in 20 crops of 41 countries. It is one of the most important weeds of cereals in Argentina, Canada, Kenya, South Africa and the USA; maize in the former Soviet Union; and sugarbeet in Spain. It is a principal weed of cereals in Argentina, Australia, Canada, UK, Finland, New Zealand, Tanzania, and the USA; maize in Italy; flax in Australia, Brazil, Canada, and the USA; potatoes in Chile; sugarbeet in former Czechoslovakia, UK and Germany; beans in England; vegetables in Argentina, Bulgaria, Chile, UK and New Zealand; peas in Bulgaria and New Zealand; onions in Argentina and UK; and sorghum in Italy.
By climbing up the crop, F. convolvulus causes lodging in grain crops (Neururer, 1961; Hume et al., 1983), and can cause harvesting problems when its vines wrap around moving parts of machinery (Forsberg and Best, 1964; Fabricius and Nalewaja, 1968).
In addition, high weed densities can raise the moisture content of harvested grain (Neururer, 1961) and contribute to heating in storage when harvested with cereals as a seed contaminant (Holm et al., 1991). F. convolvulus produces large amounts of seed and they are often difficult to separate from grain crops, because of their similar size. Therefore, F. convolvulus is a serious contaminant of seed stocks in several places in the world (Gooch, 1963; Bogdan, 1965).
The general importance of this weed is due to its ability to emerge throughout the growing season. Forsberg and Best (1964) showed that late-emerging seedlings are likely to escape herbicide spraying and could be a potential source of re-infestations, especially where competition from other weeds has been eliminated.
Furthermore, F. convolvulus may serve as an alternate host for disease organisms affecting crops (Cooper and Harrison, 1973; Bendixen et al., 1979).
According to Holm et al. (1991), F. convolvulus is ranked as a principal or serious weed in 20 crops of 41 countries. It is one of the most important weeds of cereals in Argentina, Canada, Kenya, South Africa and the USA; maize in the former Soviet Union; and sugarbeet in Spain. It is a principal weed of cereals in Argentina, Australia, Canada, UK, Finland, New Zealand, Tanzania, and the USA; maize in Italy; flax in Australia, Brazil, Canada, and the USA; potatoes in Chile; sugarbeet in former Czechoslovakia, UK and Germany; beans in England; vegetables in Argentina, Bulgaria, Chile, UK and New Zealand; peas in Bulgaria and New Zealand; onions in Argentina and UK; and sorghum in Italy.
Risk and Impact Factors
Invasiveness
Proved invasive outside its native range
Has a broad native range
Abundant in its native range
Highly adaptable to different environments
Is a habitat generalist
Tolerates, or benefits from, cultivation, browsing pressure, mutilation, fire etc
Pioneering in disturbed areas
Tolerant of shade
Highly mobile locally
Benefits from human association (i.e. it is a human commensal)
Fast growing
Has propagules that can remain viable for more than one year
Impact outcomes
Damaged ecosystem services
Ecosystem change/ habitat alteration
Modification of successional patterns
Monoculture formation
Negatively impacts agriculture
Threat to/ loss of native species
Impact mechanisms
Competition - monopolizing resources
Competition - smothering
Pest and disease transmission
Rapid growth
Likelihood of entry/control
Highly likely to be transported internationally accidentally
Difficult to identify/detect as a commodity contaminant
Difficult to identify/detect in the field
Difficult/costly to control
Uses
Tkachuk and Mellish (1977) stated that although being comparatively low in protein, oil and fibre content, seeds of F. convolvulus may be promising as a nutritious food or feed, because its amino acid composition is similar to that of cultivated buckwheat. The high lysine content of the seeds makes them a reasonable supplement to feeds otherwise consisting largely of cereals.
The seeds are supposed to have been used as food or feed in neolithic to medieval times (Eggers, 1979; Willerding, 1981; Hanf, 1990), but are too small and low-yielding to be grown commercially today. According to Boatman (1987), F. convolvulus is a host plant for insects eaten by gamebird chicks in the UK, for example, the grey partridge (Perdix perdix).
The seeds are supposed to have been used as food or feed in neolithic to medieval times (Eggers, 1979; Willerding, 1981; Hanf, 1990), but are too small and low-yielding to be grown commercially today. According to Boatman (1987), F. convolvulus is a host plant for insects eaten by gamebird chicks in the UK, for example, the grey partridge (Perdix perdix).
Prevention and Control
Due to the variable regulations around (de)registration of pesticides, your national list of registered pesticides or relevant authority should be consulted to determine which products are legally allowed for use in your country when considering chemical control. Pesticides should always be used in a lawful manner, consistent with the product's label.
Cultural Control
According to Holm et al. (1991), tillage implements and rotations have only limited success in controlling F. convolvulus because of its large seedbank, the ability of seeds to emerge from great depth, the persistent emergence of new seedlings throughout the growing season and the large quantities of seeds produced by the plants.
Nevertheless, there are several methods suggested which could be used to reduce possible yield losses caused by this weed. Koch (1964) observed a stimulation in emergence of F. convolvulus by harrowing, which could be taken advantage of in the control of this weed in cereal cultivation. Gruenhagen and Nalewaja (1969) and Messersmith and Nalewaja (1969) stated that higher seeding rates of the crop result in a reduction of relative yield losses through competition of F. convolvulus in wheat or flax. Moreover, Smith (1980) found out that a significant reduction of F. convolvulus could be attained by discing after barley harvest.
Different results have been published regarding the effects of tillage techniques as a means for weed control. Thompson et al. (1984) and Dessaint et al. (1993) concluded that no-tillage cultivation or reduced tillage resulted in increasing densities of F. convolvulus in comparison to conventional tillage, whereas Hoffman-Kakol et al. (1982) observed a reduction in the relative abundance of F. convolvulus in directly drilled maize after rye. Nielsen and Pinnerup (1982) also reported a decline in F. convolvulus populations caused by reduced cultivation in spring barley.
Although effective for the control of other weeds, harrowing in the dark showed no significant influence on emergence of F. convolvulus seedlings in comparison to daylight harrowing in Swedish field trials (Ascard, 1992).
According to Rajczyova (1978), monocultures of winter wheat and spring barley lead to an increase in F. convolvulus incidence.
Biological Control
Adkins and Sowerby (1996) revealed that the weed Parthenium hysterophorus has allelopathic potential against P. covolvulus. They used leachate from the leaves of P. hysterophorus to depress germination and seedling growth of F. convolvulus and several other weeds.
In a search for potential biological control agents for weeds, species of Botrytis were isolated from seedlings of F. convolvulus infected with pathogenic fungi in Canada (Mortensen and Molloy, 1993). In Argentina, the fungus Puccinia polygoni-amphibii was found to cause sufficient damage to warrant investigation as a biocontrol agent against F. convolvulus (Dal-Bello and Carranza, 1995).
Chemical Control
A considerable number of selective herbicides sprayed alone, in mixtures either pre- or post-emergence have been used to control F. convolvulus: aclonifen, atrazine, aziprotryne, benfluralin, benazolin, bentazone, bifenox, bromoxynil, chloramben, chlorbromuron, chloridazon, chlorsulfuron, chlorthal-dimethyl, clopyralid, cyanazine, cycloate, 2,4-D, desmedipham, dicamba, dichlorprop, dinitramine, diphenamid, ethalfluralin, ethofumesate, fluorochloridone, fluroxypyr, glufosinate-ammonium, haloxyfop, ioxynil, isopropalin, linuron, lenacil, MCPA, MCPB, mecoprop, metamitron, metolachlor, metobromuron, metsulfuron, metribuzin, napropamide, oryzalin, oxadiazon, oxyfluorfen, pendimethalin, phenmedipham, picloram, prometryn, propachlor, propyzamide, pyridate, sethoxydim, thifensulfuron, triasulfuron, tribenuron, trifluralin and triflusulfuron (Haas and Streibig 1982; Anderson et al., 1996; Fain, 1986; Nohl-Weiler and Hindersmann, 1986; Pimpini et al., 1986; Shalna and Melamed, 1986; Drazic et al., 1987; Kapros et al., 1988; Prochazka et al., 1988; Klingaman and Peeper, 1989; Milusher et al., 1989; Rapparini et al., 1989; Arends and Pegg, 1990; Lalova and Bogdanovske, 1990; Labza et al., 1990; Drazic and Glusac, 1991; Sysmans et al., 1991; Muller, 1992; Gvozdenovic-Varga et al., 1992; Meinlschmidt and Karch, 1992; Hallgren, 1993; Mitchell and Abernethy, 1993; Andersson, 1994; Rapparini, 1995, 1996; Bouma et al., 1996; Fields et al., 1996; Campagna and Rapparini, 1997; Toth and Peter, 1997).
Herbicide resistence has been reported for the following active ingredients: chlorsulfuron in Australia (Adkins et al., 1997); 2,4-D in Lithuania, Hungary and China (Aleksinas, 1984; Nemeth, 1985; Tu, 1989), 2,4-TB in Czechoslovakia (Bojas, 1987); MCPA in the former USSR and Hungary (Ryzhaya et al. 1984; Nemeth, 1985); metolachlor in the former USSR (Zuza, 1983; Veselovskii and Saulyak, 1984); metribuzin in Poland (Sawicka and Skalski, 1996); quinmerac in the UK (Boatman, 1991); simazine (Stryckers and van Himme, 1974); terbacil in Hungary (Nagy et al., 1978); and triazines in Czechoslovakia, Germany and the Netherlands (Valkova, 1975; Kees, 1988; van Oorschot, 1989).
Skorobogatova and Mirchinik (1985) revealed that the microbial metabolite citrin applied in wheat fields could decrease F. convolvulus incidence markedly, with negligible effects on wheat growth and yield. Gleason and Case (1986) showed that the algicide cyanobacterin inhibited the growth of F. convolvulus when sprayed onto the leaves of 1- to 2-week-old plants. Bryan et al. (1995) reported that a monic acid derivative of pseudomonic acid (derived from compounds produced by Pseudomonas fluorescens) was active against a range of broadleaved weeds, including F. convolvulus, in field trials performed in Canada. Barley was not affected by the monic acid derivative, although unacceptable damage to wheat did occur.
According to Holm et al. (1991), tillage implements and rotations have only limited success in controlling F. convolvulus because of its large seedbank, the ability of seeds to emerge from great depth, the persistent emergence of new seedlings throughout the growing season and the large quantities of seeds produced by the plants.
Nevertheless, there are several methods suggested which could be used to reduce possible yield losses caused by this weed. Koch (1964) observed a stimulation in emergence of F. convolvulus by harrowing, which could be taken advantage of in the control of this weed in cereal cultivation. Gruenhagen and Nalewaja (1969) and Messersmith and Nalewaja (1969) stated that higher seeding rates of the crop result in a reduction of relative yield losses through competition of F. convolvulus in wheat or flax. Moreover, Smith (1980) found out that a significant reduction of F. convolvulus could be attained by discing after barley harvest.
Different results have been published regarding the effects of tillage techniques as a means for weed control. Thompson et al. (1984) and Dessaint et al. (1993) concluded that no-tillage cultivation or reduced tillage resulted in increasing densities of F. convolvulus in comparison to conventional tillage, whereas Hoffman-Kakol et al. (1982) observed a reduction in the relative abundance of F. convolvulus in directly drilled maize after rye. Nielsen and Pinnerup (1982) also reported a decline in F. convolvulus populations caused by reduced cultivation in spring barley.
Although effective for the control of other weeds, harrowing in the dark showed no significant influence on emergence of F. convolvulus seedlings in comparison to daylight harrowing in Swedish field trials (Ascard, 1992).
According to Rajczyova (1978), monocultures of winter wheat and spring barley lead to an increase in F. convolvulus incidence.
Biological Control
Adkins and Sowerby (1996) revealed that the weed Parthenium hysterophorus has allelopathic potential against P. covolvulus. They used leachate from the leaves of P. hysterophorus to depress germination and seedling growth of F. convolvulus and several other weeds.
In a search for potential biological control agents for weeds, species of Botrytis were isolated from seedlings of F. convolvulus infected with pathogenic fungi in Canada (Mortensen and Molloy, 1993). In Argentina, the fungus Puccinia polygoni-amphibii was found to cause sufficient damage to warrant investigation as a biocontrol agent against F. convolvulus (Dal-Bello and Carranza, 1995).
Chemical Control
A considerable number of selective herbicides sprayed alone, in mixtures either pre- or post-emergence have been used to control F. convolvulus: aclonifen, atrazine, aziprotryne, benfluralin, benazolin, bentazone, bifenox, bromoxynil, chloramben, chlorbromuron, chloridazon, chlorsulfuron, chlorthal-dimethyl, clopyralid, cyanazine, cycloate, 2,4-D, desmedipham, dicamba, dichlorprop, dinitramine, diphenamid, ethalfluralin, ethofumesate, fluorochloridone, fluroxypyr, glufosinate-ammonium, haloxyfop, ioxynil, isopropalin, linuron, lenacil, MCPA, MCPB, mecoprop, metamitron, metolachlor, metobromuron, metsulfuron, metribuzin, napropamide, oryzalin, oxadiazon, oxyfluorfen, pendimethalin, phenmedipham, picloram, prometryn, propachlor, propyzamide, pyridate, sethoxydim, thifensulfuron, triasulfuron, tribenuron, trifluralin and triflusulfuron (Haas and Streibig 1982; Anderson et al., 1996; Fain, 1986; Nohl-Weiler and Hindersmann, 1986; Pimpini et al., 1986; Shalna and Melamed, 1986; Drazic et al., 1987; Kapros et al., 1988; Prochazka et al., 1988; Klingaman and Peeper, 1989; Milusher et al., 1989; Rapparini et al., 1989; Arends and Pegg, 1990; Lalova and Bogdanovske, 1990; Labza et al., 1990; Drazic and Glusac, 1991; Sysmans et al., 1991; Muller, 1992; Gvozdenovic-Varga et al., 1992; Meinlschmidt and Karch, 1992; Hallgren, 1993; Mitchell and Abernethy, 1993; Andersson, 1994; Rapparini, 1995, 1996; Bouma et al., 1996; Fields et al., 1996; Campagna and Rapparini, 1997; Toth and Peter, 1997).
Herbicide resistence has been reported for the following active ingredients: chlorsulfuron in Australia (Adkins et al., 1997); 2,4-D in Lithuania, Hungary and China (Aleksinas, 1984; Nemeth, 1985; Tu, 1989), 2,4-TB in Czechoslovakia (Bojas, 1987); MCPA in the former USSR and Hungary (Ryzhaya et al. 1984; Nemeth, 1985); metolachlor in the former USSR (Zuza, 1983; Veselovskii and Saulyak, 1984); metribuzin in Poland (Sawicka and Skalski, 1996); quinmerac in the UK (Boatman, 1991); simazine (Stryckers and van Himme, 1974); terbacil in Hungary (Nagy et al., 1978); and triazines in Czechoslovakia, Germany and the Netherlands (Valkova, 1975; Kees, 1988; van Oorschot, 1989).
Skorobogatova and Mirchinik (1985) revealed that the microbial metabolite citrin applied in wheat fields could decrease F. convolvulus incidence markedly, with negligible effects on wheat growth and yield. Gleason and Case (1986) showed that the algicide cyanobacterin inhibited the growth of F. convolvulus when sprayed onto the leaves of 1- to 2-week-old plants. Bryan et al. (1995) reported that a monic acid derivative of pseudomonic acid (derived from compounds produced by Pseudomonas fluorescens) was active against a range of broadleaved weeds, including F. convolvulus, in field trials performed in Canada. Barley was not affected by the monic acid derivative, although unacceptable damage to wheat did occur.
Links to Websites
Name | URL | Comment |
---|---|---|
GISD/IASPMR: Invasive Alien Species Pathway Management Resource and DAISIE European Invasive Alien Species Gateway | https://doi.org/10.5061/dryad.m93f6 | Data source for updated system data added to species habitat list. |
Global register of Introduced and Invasive species (GRIIS) | http://griis.org/ | Data source for updated system data added to species habitat list. |
References
Acevedo-Rodríguez P, Strong MT, 2012. Catalogue of the Seed Plants of the West Indies. Smithsonian Contributions to Botany, 98:1192 pp. Washington DC, USA: Smithsonian Institution. http://botany.si.edu/Antilles/WestIndies/catalog.htm
Adkins SW, Sowerby MS, 1996. Allelopathic potential of the weed, Parthenium hysterophorus L., in Australia. Plant Protection Quarterly, 11(1):20-23; 16 ref.
Adkins SW, Wills D, Boersma M, Walker SR, Robinson G, McLeod RJ, Einan JP, 1997. Weed resistance to chlorsulfuron and atrazine from the north-east grain region of Australia. Weed Research, 37(5):343-349.
Aleksinas AP, 1984. Application of herbicides to winter wheat. Khimiya v Sel'skom Khozyaistve, 22(1):39-41.
Alex JF, 1982. Canada. In: Holzner W, Numata M, eds. Biology and Ecology of Weeds. The Hague, Netherlands: Dr. W. Junk Publishers, 309-331.
Anderson MD, Staska KJ, Mayland PG, 1986. Grass and broadleaf weed control in cereals with tank-mixes of diclofop, bromoxynil and MCPA ester. Proceedings of the Western Society of Weed Science., Vol.39:33-34.
Andersson L, 1994. Effects of MCPA and tribenuron-methyl on seed production and seed size of annual weeds. Swedish Journal of Agricultural Research, 24(2):49-56
Andreasen C, Streibig JC, 1990. Impact of soil factors on weeds in Danish cereals crops. Symposium on integrated weed management in cereals. Proceedings of an EWRS Symposium. Wageningen, Netherlands: European Weed Research Society, 53-59.
Andreasen C, Streibig JC, Haas H, 1991. Soil properties affecting the distribution of 37 weed species in Danish fields. Weed Research, 31:181-187.
Andreasen C, Stryhn H, Streibig JC, 1996. Decline of the flora in Danish arable fields. Journal of Applied Ecology, 33(3):619-626; 44 ref.
Arends L, Pegg IR, 1990. Thifensulfuron methyl with metsulfuron methyl - a new sulfonylurea herbicide for broad-leaved weed control in winter cereals in New South Wales and Queensland. Proceedings of the 9th Australian Weeds Conference, 60-64
Ascard J, 1992. Harrowing at night - influence on the emergence of weeds. Swedish Crop Protection Conference. Weeds and weed control. Uppsala, Sweden: Swedish University of Agricultural Sciences, 33:163-170.
Bain OG, Johnson JL, 1986. Results of an experimental use program in the northwest for Assert (AC 222, 93) in small grains. Proceedings of the Western Society of Weed Science, 39:158-159.
Bakos J, Eifert G, Bihari F, Nagy M, 1991. HC-252 - a new selective herbicide for the post-emergence control of dicotyledonous weeds. Proceedings of the Brighton Crop Protection Conference, Weeds, 1:83-86.
Banaras M, 1993. Impact of weed competition on potato production. Pakistan Journal of Agricultural Research, 14 (1):64-71.
Bazdyrev GI, Kupryushkin VA, Lyutov VN, Ontpv AKh, 1984. Effect of mineral fertilizers on specific composition of weeds in cereals. Nauchnye osnovy povysheniya effektivnosti udobrenii v Nechernozemnoi zone Moscow, USSR, 78-84
Bendixen LE, Reynolds DA, Riedel RM, 1979. An annotated bibliography of weeds as reservoirs for organisms affecting crops. 1. Nematodes. Research Bulletin No. 1109. Ohio, USA: Ohio Agricultural Research and Development Center.
Berezkin Yu N, Skorolupova TP, 1987. Dicotyledonous annual weeds. Zashchita Rastenii, Moscow, 7:22-23.
Blackshaw RE, Larney FO, Lindwall CW, Kozub GC, 1994. Crop rotation and tillage effects on weed populations on the semi-arid Canadian prairies. Weed Technology, 8(2):231-237
Boatman ND, 1987. Selective grass weed control in cereal headlands to encourage game and wildlife. Proceedings 1987 British Crop Protection Conference, Weeds., Vol. 1:277-284; 21 ref.
Boatman ND, 1991. Selective control of cleavers (Galium aparine) in conservation headlands with quinmerac. In: Proceedings of the 1991 Brighton Crop Protection Conference - Weeds. Farnham, UK: British Crop Protection Council, 2:669-676.
Bogdan AV, 1965. Weeds in Kenya wheat. Weed Research, 5:351-352.
Bojas Z, 1987. New information on weed control in maize undersown with alfalfa and red clover. Agrochemia, 27(2):55-59
Borgo A, 1980. Comparison of the efficacy and selectivity of chemical treatments to control Polygonum convolvulus in wheat. Resumos XIII Congresso Brasileiro de Herbicidas e Ervas Daninhas, Bahia, 1980, 46
Borowiec S, Kuszelewski L, Kutyna I, LeSnik T, lukaszuk T, 1985. The effect of long-term fertilizer use on soil properties and weed infestation of rye, spring barley, Sinapis alba and potato crops. Part II. The effect of different fertilizer treatments on weed infestation of individual crops. Zeszyty Naukowe Akademii Rolniczej w Szczecinie, Rolnictwo, No.37:71-86.
Borowiec S, Kutyna I, 1988. Effect of mineral and organic (farm manure and cattle slurry) fertilization and their interaction on weedy state of cultivated plants. Zeszyty Naukowe - Akademii Rolniczej w Szczecinie, Rolnictwo, No. 45:3-26.
Boulet C, Hammoumi M, 1984. Distribution of the main weeds in the oceanic region of western Morocco. Comptes Rendu du 7eme Colloque International sur l'Ecologie, la Biologie et la Systematique des Mauvaises Herbes. Paris, France: COLUMA/EWRS, 1:173-182.
Bouma E, Weide RYvan der, Floot HWG, 1996. Influence of weather parameters on efficacy of reduced dosages of herbicides in winter wheat. Bulletin OEPP, 26(3/4):651-657; 12 ref.
Brendler F, 1995. Specific and effective weed control. Kartoffelbau, 46(4):172-173.
BrSutigam H, 1995. Does nitrogen fertilizer affect weed populations in sugarbeet? PSP Pflanzenschutz Praxis, 1:24-26.
Bryan IB, Rice MJ, Bartley MR, Jutsum AR, Pastushok G, 1995. A monic acid derivative: evaluation as a cereal herbicide. Brighton crop protection conference: weeds. Proceedings of an international conference, Brighton, UK, 20-23 November 1995., Vol. 2:725-730; 4 ref.
Bujßn M, Castelao AM, Sainz MJ, 1995. Weeds in an ecological culture of asparagus (Asparagus officinalis L.) in Galicia: first results. Proceedings of the 1995 Congress of the Spanish Weed Science Society, Huesca, Spain, 14-16 November 1995., 83-86; 7 ref.
Caballero R, Barro C, Alzueta C, Arauzo M, Hernaiz PJ, 1995. Weed control and herbicide tolerance in a common vetch-oat intercrop. Weed Science, 43(2):283-287
Cadar T, 1983. Dynamics of weed seeds in the sowing material in Maramures district during 1970-1981. Probleme de Agrofitotehnie Teoretica si Aplicata, 5(4):395-411; 6 ref.
Campagna G, Rapparini G, 1997. Fallopia convolvulus (L.) Holub., black bindweed (code: POLCO or FALCO). Informatore Agrario, 53(10):92.
Catullo JC, Rodriguez ML, Sosa CA, Colombo I, 1983. Determination of the critical period of weed competition in sunflowers. Malezas, 11(4):150-164
Chancellor RJ, 1964. The depth of weed seed germination in the field. Proceedings of the 7th British Weed Control Conference, 2:607-613.
Chen TB, Lin C, 1989. Phytocoenological features and control strategies of weeds. Proceedings, 12th Asian-Pacific Weed Science Society Conference., No. 1:73-78.
Chepil W, 1946. Germination of weed seeds: I. Longevity, periodicity of germination and vitality of seeds in cultivated soil. Scientific Agriculture, 26:307-346.
Chippendale HG, Milton WEJ, 1934. On the viable seeds present in the soil beneath pastures. Journal of Ecology, 22:508-531.
Chirita N, 1990. Usefulness of some treatments with different herbicides in mono- and dicotyledonous weed control in oilseed flax crops. Cercetari Agronomice în Moldova, 23(1):53-55; 4 ref.
Clapham AR, Tutin TG, Warburg EF, 1962. Flora of the British Isles. Second edition. Cambridge, UK: Cambridge University Press.
Conert HJ, Hamann U, Schulte-Motel W, Wagenitz G, 1981. Gustav Hegi: Illustrierte Flora von Mitteleuropa Band III, Teil 1. Third edition. Berlin, Hamburg: Verlag Paul Parey.
Conn JS, Deck RE, 1995. Seed viability and dormancy of 17 weed species after 9.7 years of burial in Alaska. Weed Science, 43(4):583-585; 10 ref.
Conners IL, 1967. An annotated index of plant diseases in Canada and fungi recorded on plants in Alaska, Canada and Greenland. Research Branch, Canada Department of Agriculture, Publ. 1251.
Cooper JI, Harrison BD, 1973. The role of weed hosts and the distribution and activity of vector nematodes in the ecology of tobacco rattle virus. Annals of Applied Biology, 73(1):53-66
Currie RS, Peeper TF, 1986. Effect of harvest method on weed seed germination and potential emergence of resistance. Proceedings of the 39th Annual Meeting of the Southern Weed Science Society, 395.
Dahl GK, Busse SR, McNeil WK, 1986. Imazamethabenz: results of 1986 field trials. Proceedings, North Central Weed Control Conference, Milwaukee, Wisconsin, USA, 41:62.
DAISIE, 2015. Delivering Alien Invasive Species Inventories for Europe. European Invasive Alien Species Gateway. www.europe-aliens.org/default.do
Dal Bello GM, Carranza MR, 1995. Weed diseases in La Plata area II. Identification of pathogens with potential for weed biocontrol programmes. Revista de la Facultad de Agronomi^acute~a (La Plata), 71(1):7-14; 21 ref.
Dalmau L, Plana E, Verdu AM, 1993. Solarization, tillage and weed control in Valles Oriental (Barcelona). Proceedings of the 1993 Congress of the Spanish Weed Science Society. Madrid, Spain: Sociedad Espanola de Malherbologia, 264-267.
Dechkov Z, 1987. The problem of weed infestation in the Pleven district. Pochvoznanie, Agrokhimiya i Rastitelna Zashchita, 22(3):118-124; 6 ref.
Dechkov Z, 1989. Potential weed infestation in the south-west part of Bulgaria. Rasteniev"dni Nauki, 26(4):122-126.
Decraene LPR, Hong SukPyo, Smets E, 2000. Systematic significance of fruit morphology and anatomy in tribes Persicarieae and Polygoneae (Polygonaceae). Botanical Journal of the Linnean Society [Under the microscope: plant anatomy and systematics. Proceedings of a symposium held in honour of David Cutler at the Linnean Society of London, UK, 9-10 September 1999, co-sponsored by the Linnean Society and the Systematics Association.], 134(1/2):301-337.
Dessaint F, Chadoeuf R, Barralis G, 1991. Spatial pattern analysis of weed seeds in the cultivated soil seed bank. Journal of Applied Ecology, 28(2):721-730
Dessaint F, Chadoeuf R, Barralis G, 1993. The long term influence of soil cultivation on the density of weed emergence under non-herbicide conditions. Canadian Journal of Botany, 71(7):919-926
Dochkova B, 1969. Vliyanie na khranitelnite rasteniga varkhu razvitieto i plodovitostta na grasinskata nastenka. Rastenievd Nauki, 6:127-134.
Dochkova B, 1972. Some ecological studies on Mamestra brassicae L. (Lepidoptera Noctuidae). Gradinarska i Lozarska Nauka, 9(1):77-86
Dosland JG, Arnold JD, 1966. Leaf area development and dry matter production of wheat and wild buckwheat growing in competion. Abstracts of the Meetings of the Weed Science Society of America, 56.
Drazic D, Glusac D, 1987. Herbicide application depending on the intensity of weed occurrence in wheat. Zastita Bilja, 38(4):387-395
Drazic D, Glusac D, 1991. Effectiveness and selectiveness of herbicide combination in maize. Savremena Poljoprivreda, 39(5):33-38
Drazic D, Kosovac Z, Glusac D, 1987. Effects of herbicides on wheat yields and germination of wheat seeds. Fragmenta Herbologica Jugoslavica, 16(1-2):203-208
Duer I, 1986. Chemical composition of weeds and nutrient uptake by weeds and cereals in rotations with different percentage of cereal crops. Pamietnik Pulawski, 88:191-204.
Durgan BR, Yenish JP, Daml RJ, Miller DW, 1997. Broadleaf weed control in hard red spring wheat (Triticum aestivum) with F8426. Weed Technology, 11(3):489-495; 6 ref.
Eggers T, 1979. Development and changes of field weed vegetation. Berichte der Internationalen Symposien der Internationalen Vereinigung für Vegetationskunde: Werden und Vergehen von Pflanzengesellschaften; Rinteln, 503-527.
Erviö R, Hyvarinen S, Ervio LR, Salonen J, 1994. Soil properties affecting weed distribution in spring cereal and vegetable fields. Agricultural Science in Finland, 3(5):497-504.
Fabricius, LJ, Nalewaja, D, 1968. Competition between wheat and wild buckwheat. Weed Science, 16:204 208.
Fain DM, 1986. Wild buckwheat control in wheat - 1986. Proceedings, Southern Weed Science Society, 39th Annual Meeting, 482.
Fain DM, Peeper TF, Greer HA, 1980. Wild buckwheat in Oklahoma wheat: problems and control. Proceedings of the 33rd Annual Meeting of the Southern Weed Science Society, 66
Falcon LMF de, Bedendo ED, Farias G, 1991. Sequence of weed emergence. Proceedings of the 12th Argentine meeting on weeds and their control. Buenos Aires, Argentina: Asociacion Argentina para el Control de Malezas, 1:45-52.
FAO, 2015. Polygonum convolvulus L. Online resources. http://www.fao.org/agriculture/crops/thematic-sitemap/theme/biodiversity/weeds/listweeds/pol-con/en/
Felton WL, Wicks GA, Welsby SM, 1994. A survey of fallow practices and weed floras in wheat stubble and grain sorghum in northern New South Wales. Australian Journal of Experimental Agriculture, 34(2):229-236.
Fields SC, Mireles-Lo L, Gerwick BC, 1996. Hydroxycornexistin: a new phytotoxin from Paecilomyces variotii. Journal of Natural Products, 59(7):698-700; 6 ref.
Fisher SJ, 1992. Herbicide development designed for cost effective weed control. Aspects of Applied Biology, 32:59-64.
Fleck NG, Mengarda IP, Pinto JJO, 1989. Weed interference in sunflower. Competition in space. Pesquisa Agropecuaria Brasileira, 24(9):1131-1137.
Flora of China Editorial Committee, 2015. Flora of China. St. Louis, Missouri and Cambridge, Massachusetts, USA: Missouri Botanical Garden and Harvard University Herbaria. http://www.efloras.org/flora_page.aspx?flora_id=2
Flora of Pakistan Editorial Committee, 2013. Flora of Pakistan, eFloras website. St. Louis, Missouri and Cambridge, Massachusetts, USA: Missouri Botanical Garden and Harvard University Herbaria. http://www.efloras.org/flora_page.aspx?flora_id=5
Fontana SA, 1980. Control of foxtail grass and broad-leaved weeds in barley with propanil. Proceedings 1979 North Central Weed Control Conference, 34:36.
Forcella F, Wilson RG, Dekker J, Kremer RJ, Cardina J, Anderson RL, Alm D, Renner KA, Harvey RG, Clay S, Buhler DD, 1997. Weed seed bank emergence across the Corn Belt. Weed Science, 45(1):67-76.
Forsberg DE, 1955. Gastrophysa polygoni L., a beetle that attacks wild buckwheat (Polygonum convolvulus). North Central Weed Control Conference, 12th Annual Report, 183.
Forsberg DE, Best KF, 1964. The emergence and plant development of wild buckwheat (Polygonum convolvulus L.). Canadial Journal of Plant Science, 44:100-103.
Franzini E, 1982. Italy. In: Holzner W, Numata M, eds. Biology and ecology of weeds. The Hague, Netherlands: Dr. W. Junk Publishers, 245-256.
Freeman CC, Hinds HR, 2005. 29. Fallopia Adanson, Fam. Pl. 2: 277, 557. 1763, name conserved. In: Flora of North America North of Mexico. Volume 5. Magnoliphyta: Caryophyllidae, Part 2 [ed. by Flora of North America Editorial Committee]. New York, NY, USA: Oxford University Press, 541-546.
Friesen G, Shebeski LH, 1960. Economic losses caused by weed competition in Manitoba grain fields. I. Weed species, their relative abundance and their effect on crop yields. Canadian Journal of Plant Science, 40:457-467.
Fuchs H, Voit B, 1992. Purity and contamination in cereal seed samples - long term survey from results of seed testing. Bayerisches Landwirtschaftliches Jahrbuch, 69(6):757-777
Gaskin TA, 1958. Weed hosts of Meloidogyne incognita in Indiana. Plant Disease Reporter, 42:802-803.
Gerard PJ, 1989. Influence of egg depth in host plants on parasitism of Scolypopa australis (Homoptera: Ricaniidae) by Centrodora scolypopp (Hymenoptera: Aphelinidae). New Zealand Entomologist, 12:30-34.
Gill KS, Arshad MA, 1995. Weed flora in the early growth period of spring crops under conventional, reduced, and zero tillage systems on a clay soil in northern Alberta, Canada. Soil & Tillage Research, 33(1):65-79
Gleason FK, Case DE, 1986. Activity of the natural algicide, cyanobacterin, on angiosperms. Plant Physiology, 80(4):834-837; 17 ref.
Gooch S, 1963. The occurrence of weed seeds in samples tested by the Official Seed Testing Station 1960-1961. Journal of the National Institute of Agricultural Botany, 9(3):353-371.
Goodney T, 1947. On the stem eelworm, Anguillulina dipsaci, attacking oats, onions, field beans, parsnips, rhubarb and certain weeds. Journal of Helminthology, 22:1-12.
Goodwin MS, Morrison IN, Thomas AG, 1986. A weed survey of pedigreed alfalfa seed fields in Manitoba. Canadian Journal of Plant Science, 66(2):413-416
Grigaliunaite B, Kacergius A, 1995. The fungi of weed seeds. Ekologija, No. 3:50-56; 13 ref.
Gruenhagen, RD, Nalewaja, JD, 1969. Competition between flax and wild buckwheat. Weed Science, 17:380-384.
Gvozdenovic-Varga J, Glusac D, Takac A, 1992. Applicability of herbicides in onions and their effect on yield level. Savremena Poljoprivreda, 40(1/2):196-201.
Haas H, Streibig JC, 1982. Changing patterns of weed distribution as a result of herbicide use and other agronomic factors. In: LeBaron HM, Gressel J, eds. Herbicide Resistance in Plants. New York, USA: John Wiley and Sons, 57-79.
Hallgren E, 1993. Effects of some herbicides or mixtures of herbicides on individual dicot weed species at different doses and developmental stages. Vaxtodling, Institutionen for Vaxtodling, Sveriges Lantbruksuniversitet, 44.
Hallgren E, 1996. Do the weed flora and effect of a herbicide change with time?. Proceedings of the second international weed control congress, Copenhagen, Denmark, 25-28 June 1996: Volumes 1-4., 1355-1368; 13 ref.
Hallgren E, 1996. Occurrence of broad-leaved weeds on different soils in different crops in Sweden. Swedish Journal of Agricultural Research, 26(3):115-123.
Haman CD, Peeper TF, 1983. The effect of shade on wild buckwheat. Proceedings, Southern Weed Science Society, 36th Annual Meeting, 348.
Hanf M, 1982. AckerunkrSuter Europas mit ihren Keimlingen und Samen. Ludwigshafen, Germany: BASF Aktiengesellschaf.
Hanf M, 1990. Farbatlas Feldflora - Wildkraeuter und Unkraeuter. Stuttgart, Germany: Ulmer.
Hashimoto G, 1982. Brazil. In: Holzner W, Numata M, eds. Biology and Ecology of Weeds. The Hague, Netherlands: Dr. W. Junk, 333-337.
Hilbig W, 1982. Mongolia. In: Holzner W, Numata M, eds. Biology and Ecology of Weeds. The Hague, Netherlands: Dr. W. Junk Publishers, 277-279.
Hilbig W, Mahn EG, 1971. The mapping of arable weeds as a basis for the objective use of herbicides. SYS Reporter, 3:2-23.
Himme M van, Stryckers J, Bulcke R, 1983. Fibre flax. Mededelingen van het Centrum voor Onkruidonderzoek, No.38:107-114
Hochol T, 1990. Segetal weed communities in the Lososina Valley in the Beskid Wyspowy: Part II. The weediness of cereals. Zeszyty Naukowe Akademii Rolniczej im. H. Kollataja w Krakowie, Rolictwo, 29:77-91.
Hoffman-Kakol I, Dzienia S, Stankiewicz J, 1982. The influence of simplified pre-sowing cultivation on weed infestation of maize grown as a second crop. Zeszyty Naukowe Akademii Rolniczej w Szczecinie, Rolnictwo, No.94:83-94; 13 ref.
Holm LG, Pancho JV, Herberger JP, Plucknett DL, 1979. A geographical atlas of world weeds. New York, USA: John Wiley and Sons, 391 pp.
Holm LG, Pancho JV, Herberger JP, Plucknett DL, 1991. A Geographic Atlas of World Weeds. Malabar, Florida, USA: Krieger Publishing Company.
Holm LG, Plucknett DL, Pancho JV, Herberger JP, 1977. The World's Worst Weeds. Distribution and Biology. Honolulu, Hawaii, USA: University Press of Hawaii.
Holub J, 1970. Fallopia Adans. 1763 instead of Bilderdykia Dum. 1827. Folia Geobotanica Phytotaxonomica, 6:171-177.
Hübl E, Holzner W, 1982. Iran. In: Holzner W, Numata M, eds. Biology and Ecology of Weeds. The Hague, Netherlands: Dr. W. Junk Publishers, 257-266.
Hume L, 1993. Development of equations for estimating yield losses caused by multi-species weed communities dominated by green foxtail (Setaria viridis (L.) Beauv.). Canadian Journal of Plant Science, 73(2):625-635; 18 ref.
Hume L, Martinez J, Best K, 1983. The biology of Canadian weeds. 60. Polygonum convolvulus L. Canadian Journal of Plant Science, 63(4):959-971
Hunyadi K, 1973. Weed problems of the south-western arable lands of Hungary. Jugoslovenski Simpozium o Borbi protiv Korova u Brdsko-Planinskim Podrucjima, Sarajeva, 1973, 61-66
Ilic V, Kalinovic I, 1995. Contribution to knowledge of foreign matters in stored mercantile maize seed. Acta Agronomica Ovariensis, 37(2):145-152.
Ivany JA, 1987. Chlorsulfuron use in barley and residual effect on potato and rutabaga grown in rotation. Canadian Journal of Plant Science, 67(1):337-341.
Jackson MJ, Fay PK, 1979. The weed research and extension programme in Montana. Proceedings of the Western Society of Weed Science, 32:106-109.
Jordan MJ, Nadelhoffer KJ, Fry B, 1997. Nitrogen cycling in forest and grass ecosystems irrigated with <sup(15)>N-enriched wastewater. Ecological Applications, 7(3):864-881; 52 ref.
Kang BH, Kwon YW, Lee HK, 1996. Current status and problem of exotic weeds in Korea. Import and export of agricultural products and plant quarantine. '96 International Symposium, Seoul, Korea Republic, 19 May 1996., 101-128; 30 ref.
Kapros J, Petranyi I, Szito A, 1988. New possibilities of weed control in peas. Novenyvedelem, 24(2):69-70
Karczmarczyk S, Hoffman-Kakol I, Koszanski Z, 1983. A comparison of the yields of potatoes and sugarbeet grown on light soils with irrigation and different rates of mineral fertilizers. Part II. Weed infestation. Zeszyty Naukowe Akademii Rolniczej w Szczecinie, Rolnictwo, No. 31:59-65; 5 ref.
Katis NJ, Kokinis G, Eleftherohorinos I, Jones P, 1997. Arable weeds identified as new sources of beet mosaic potyvirus in Greece. Annals of Applied Biology, 130(2):255-260; 17 ref.
Kauppila R, 1990. Conventional and organic cropping systems at Suitia. IV: weeds. Journal of Agricultural Science in Finland, 62(4):331-337
Kazantseva AS, Tuganaev VV, 1972. The species composition and distribution of weed seeds in soils of field plant communities in the Tatar ASSR. Biologicheskie Nauki, 15(11):72-74.
Kees H, 1988. The development of triazine-resistant annual weeds in Bavaria and experience with their control. Gesunde Pflanzen, 40(10):407-412, 414
Klingaman TE, Peeper TF, 1989. Weed control in winter wheat (Triticum aestivum) with chlorsulfuron and CGA 131036 and comparison of modes of action. Weed Technology, 3(3):490-496
Kneeshaw PG, Kinoshita GB, Luke N, 1983. Weed control in cereals with AC 222, 239: Canadian results. Proceedings, 1983 North Central Weed Control Conference, 76.
Koch W, 1964. UnkrautbekSmpfung durch Eggen, Hacken und Meißeln in Getreide. I. Wirkungsweise und Einsatzzeitpunkt von Egge, Hacke und Bodenmeißel. Zeitschrift für Acker und Pflanzenbau, 120:369-382.
Korkut I, Kasa M, 1979. Chemical tests against the weeds causing damage to sunflowers in the Black Sea region of Turkey. Turkey, Zirai Mucadele ve Zirai Karantina Genel Mudurlugu, Arastirma Dairesi Baskanligi: Plant protection research annual (report).: Zirai mucadele arastirma yilligi., 150, 162
Korsmo E, 1954. Anatomy of weeds. Oslo, Norway: Grohndal & Sons, 172-175.
Kryukova EA, Persidskaya LT, 1986. The formation of entomofauna and pathogenic microflora in a sylvagrarian countryside: ways of improving the resistance of agricultural plant communities. Vestnik Sel'skokhozyaistvennoi Nauki, No. 4:61-66; 6 ref.
Kudsk P, Mathiassen SK, Noye G, 1997. Phenmedipham og triallat i spinat til fro. SP-Rapport Statens Planteavlsforsog, 7:139-148.
Kutschera L, 1960. Wurzelatlas mitteleuropSischer AckerunkrSuter und Kulturpflanzen. Frankfurt, Germany: DLG-Verlag.
Labza T, Stupnicka-Rodzynkiewicz E, Hochol T, Lepiarczyk A, 1990. The usage of Benomeks and Chwastoben in the control of dicotyledonous weeds growing in spring barley. Zeszyty Naukowe Akademii Rolniczej im. H. Kollataja w Krakowie, Rolictwo, 29:105-114.
Lalova M, Bogdanovske V, 1990. Weed control in winter wheat with chlorsulfuron and Lactofol O. Symposium on integrated weed management in cereals. Proceedings of an EWRS Symposium. Wageningen, Netherlands: European Weed Research Society, 415-420.
Lamb Frye AS, Kron KA, 2003. rbcL phylogeny and character evolution in Polygonaceae. Systematic Botany, 28(2):326-332.
Larsson M, 1994. Pathogenicity, morphology and isozyme variability among isolates of Aphanomyces spp. from weeds and various crop plants. Mycological Research, 98(2):231-240
Lauer E, 1953. Uber die Keimtemperatur von Ackerunkrautern und deren einfluss auf die zusammensetzung von Unkrautgesellschaften. Flora Oder Allgemeine Botanische Zeitung, 140:551-595.
MacKee HS, 1994. Catalogue of introduced and cultivated plants in New Caledonia. (Catalogue des plantes introduites et cultivées en Nouvelle-Calédonie.) Paris, France: Muséum National d'Histoire Naturelle, unpaginated.
Maharajan M, Rajendran A, 2014. Taxonomic studies on selected species of the genus Polygonum L. (Polygonaceae) in South India. Journal of Science, 4:144-148. http://www.journalofscience.net/File_Folder/144-148.pdf
Mahn EG, 1984. The influence of different nitrogen levels on the productivity and structural changes of weed communities in agro-ecosystems. Comptes Rendus du 7eme Colloque International sur l'Ecologie, la Biologie et la Systematique des mauvaises herbes. Paris, France: COLUMA/EWRS, 1:421-429.
Mayor JP, Maillard A, 1995. Results from an over-20-years-old ploughless tillage experiment at Changins. IV. Seed bank and weed control. Revue Suisse d'Agriculture, 27(4):229-236
McGinley MA, Tilman D, 1993. Short-term response of old-field plant communities to fire and disturbance. American Midland Naturalist, 129(2):409-413; 21 ref.
Meinlschmidt E, Karch K, 1992. Possibilities for suppression of weed growth with repeated application of low doses of herbicides. Zeitschrift für Pflanzenkrankheiten und Pflanzenschutz, Sonderheft, 13:529-538.
Meisel K, 1979. Changes in the segetal vegetation on the Weser marshlands at Stolzenau since 1945. Phytocoenologia, 6(10):118-130
Melo E, 2015. Polygonaceae in Lista de Espécies da Flora do Brasil (Polygonaceae in the list of species of the flora of Brazil). Rio de Janeiro, Brazil: Jardim Botânico do Rio de Janeiro. http://floradobrasil.jbrj.gov.br/jabot/floradobrasil/FB196
Messersmith CG, Nalewaja JD, 1969. Competition between wheat and wild buckwheat. Meeting of the Weed Science Society of America, Abstract No. 15.
Michal J, 1991. Weed flora of fields along the Luznice River upstream. Acta Scientifica, Vysoka Skola Zemedelska v Praze Fakulta Agronomicka v Ceskych Budejovicich, 38:69-76.
Miller SD, Alley HP, 1985. Broadleaf weed control and rotational crop response with clopyralid. Proceedings, North Central Weed Control Conference St. Louis, Missouri, USA, 40:17.
Milusher V, Balinova A, V"lkov G, 1989. Weed damage to seed potatoes. Rasteniev'dni Nauki, 26(6): 99-105.
Mitchell RB, Abernethy RJ, 1993. Tolerance of clary sage, coriander and caraway to herbicides applied pre- and post-emergence. Proceedings of the Forty Sixth New Zealand Plant Protection Conference. Rotorua, New Zealand: New Zealand Plant Protection Society, 24-29.
Mitich LW, 1975. Picloram plus 2,4-D on small grains in North Dakota. Down to Earth, 31(1):23-25
Monstvilaite J, Petroviene I, Tyla G, 1996. Tests of herbicide efficiency in different varieties of barley and oats. Lietuvos Zemdirbystes Instituto Mokslo Darbai, Zemdirbyste, No. 55:119-129; 12 ref.
Mortensen K, Molloy MM, 1993. Survey for seed-borne diseases on weed species from screening samples obtained from seed cleaning plants across Canada in 1987/88. Canadian Plant Disease Survey, 73(2):129-136; 24 ref.
Muller K, 1992. Harmony - several years' experience with the control of weeds in maize. Zeitschrift für Pflanzenkrankheiten und Pflanzenschutz, Sonderheft 13:677-681.
Mulligan GA, 1957. Chromosome numbers of Canadian weeds. Canadian Journal of Botany, 35:779-789.
Mulligan GA, 1960. Polyploidy in Canadian weeds. Canadian Journal of Genetic Cytology, 2:150-161.
Mulligan GA, Findlay JN, 1970. Reproductive systems and colonization in Canadian weeds. Canadian Journal of Botany, 48(5):859-860.
Nagy F, Foldesi D, Szalay P, 1978. Recent results in the chemical weed control of peppermint (Mentha piperita L.). Herba Hungarica, 17(1):65-81
Nakoneshny W, Friesen G, 1961. The influence of a commercial fertilizer treatment on weed competition in spring sown wheat. Canadian Journal of Plant Science, 41:231-238.
Nati D, 1994. Weed control in wheat. Terra e Sole, 49(625):426-428
Nemeth I, 1985. Weed flora of vineyards. Novenyvedelem, 21(6):252-257
Neururer H, 1961. UnkrSuter, die den MShdrusch stören. Der Pflanzenarzt, 14:61-63.
Neururer H, 1990. Experiences in the use of expanding harrows (flexible harrows) for weed control. Pflanzenschutz (Wien), 5:6-7.
Nielsen HJ, Pinnerup SP, 1982. Reduced cultivation and weeds. Weeds and Weed Control. 23rd Swedish Weed Conference, Uppsala, 1982 Sveriges Lantbruksuniversitet Uppsala Sweden, 2:370-384
Nohl-Weiler C, Hindersmann U, 1986. Several years of experience with the herbicide Racer in potatoes. Mitteilungen aus der Biologischen Bundesanstalt für Land- und Forstwirtschaft, Berlin-Dahlem, 232:350-351.
Nowicka B, 1994. Response of some winter wheat cultivars to herbicides. Ochrona Roslin, 38(5):18-20
Oorschot JLP van, 1989. Chloroplastic resistance of weeds to triazines in the Netherlands until 1988. In: Cavalloro R, Noye G, eds. Importance and perspectives on herbicide-resistant weeds. Proceedings of a meeting of the EC Experts' Group. Luxembourg: Office for Official Publications of the European Community, No. EUR 11561 EN:41-45.
Oostenbrink M, 1955. Nematologische waarnemingen. II. Aphelenchoides fragariae (Ritzema Bos, 1891) Christie, 1932 in Lilium regale Wils,. L. henryi Bak., L. sulphurgale Wallace en in de bollen van L. pumilum D.C. Verslagen en Mededelingen van de Plantenziektenkundige Dienst te Wageningen, 127:235-237.
Orchard AE, 1994. Flora of Australia. Vol. 49, Oceanic islands 1. Canberra, Australia: Australian Government Publishing Service.
Paatela J, Erviö L, 1971. Weed seeds in cultivated soils in Finland. Ann. Agric. Finnae, 10:144-152.
Pammel LH, 1913. The Weed Flora of Iowa. Des Moines, USA: Iowa Geological Society.
Petersen HI, 1960. Ukrudtsplanter og Ukrudtsbekæmpelse. K¢benhavn: Landhusholdningsselskabets Forlag.
Petunova AA, 1995. Controlling weeds resistant to 2,4-D in Russian cereal crops. Resistant Pest Management, 7(2):23.
Phillipson A, Cox TW, Elliott JG, 1972. A survey of cereal husbandry and weed control in three regions of England, 1969 and 1970. Technical Report, Agricultural Research Council, Weed Research Organization, 20.
Pimpini F, Berti A, Zanin G, 1986. Weed control in direct-drilled tomatoes. Comptes Rendus de la 13e Conference du COLUMA, Tome 3:165-175
Popay AI, Ivens GW, 1982. East Africa. In: Holzner W, Numata M, eds. Biology and Ecology of Weeds. The Hague, Netherlands: Dr. W. Junk Publishers, 345-372.
Potsch J, 1987. Results of a random sampling method for the detection and evaluation of extensive changes in the weed vegetation. Hercynia, 24(4):404-407
Prochazka I, Dvorak J, Prochazka J, 1988. Comparison of the herbicides Benazalox and Labrax in winter rape. Agrochemia, 28(2):54-57
Pulcher-HSussling M, Hurle K, 1986. Effect of N-fertilization on competition between weeds and winter wheat. Proceedings, EWRS symposium on economic weed control. Wageningen, Netherlands: EWRS, 137-144.
Rajczyova M, 1978. Causes of overproduction of weeds in cereals. Acta Botanica Slovaca Academiae Scientiarum Slovacp, A, 3:181-187
Ransom ER, 1935. The interrelations of catalase, respiration, after-ripening, and germination in some dormant seeds of the Polygonaceae. American Journal of Botany, 22:815-825.
Rapparini G, 1995. Post-emergence treatment in sugarbeet. Informatore Agrario, 51(10):81-91
Rapparini G, 1996. Weed control in sunflower. Informatore Agrario, 52(6):99-107.
Rapparini G, Fabbri M, Bartolini D, 1989. Study of herbicide degradation times and assessment of residues by bioassay. Informatore Agrario, 45(2):97-105
Reis EM, 1982. Survey of cultivated, native and weed plants, hosts of fungi that cause root rots on winter cereals and other crops. Summa Phytopathologica, 8(1/2):134-140.
Roberts HA, Chancellor RJ, 1986. Seed banks of some arable soils in the English midlands. Weed Research, UK, 26(4):251-257
Rutledge CR, McLendon T, 1996. An Assessment of Exotic Plant Species of Rocky Mountain National Park., USA: Department of Rangeland Ecosystem Science, Colorado State University, 97 pp. http://www.npwrc.usgs.gov/resource/plants/explant/index.htm
Ryzhaya MA, Murashova NF, Yanitskaya LI, 1984. Application of Basagran M to fibre flax. Khimiya v Sel'skom Khozyaistve, 12:34-35.
Sawicka B, Skalski J, 1996. Zachwaszczenie ziemniaka w warunkach stosowania herbicydu Sencor 70 WP cz. I. Skutecznosc chwastobojcza herbicydu. Roczniki Nauk Rolniczych. Seria A, Produkcja Roslinna, 112(1/2):169-182.
Schlotter P, Schuster S, 1992. Use of Starane in maize. Zeitschrift für Pflanzenkrankheiten und Pflanzenschutz, Sonderheft, 13:683-687
Schultz GE, Tichota JM, 1981. Factors influencing efficacy of low rates of glyphosate plus dicamba in reduced tillage systems. Proceedings North Central Weed Control Conference, 1981, 6:56.
Schweizer EE, Zimdahl RL, 1979. Changes in the number of weed seeds in irrigated soil under two management systems. Proceedings of the Western Society of Weed Science, 32:74.
SchwSr CH, Feyerabend G, Golz H, 1970. 100 wichtige AckerunkrSuter. Jena, Germany: Fischer.
Scott RC, Peeper TF, 1994. Economic returns from broadleaf weed control in hard red winter wheat (Triticum aestivum). Weed Technology, 8(4):797-806
Scragg EB, 1974. Regional weed problems - dicotyledonous weeds in tillage crops. Weed Control in the Northern Environment; Proceedings of a 1974 Symposium. British Crop Protection Monograph, 9:19-32.
Sgattoni P, Villani P, Ticchiati V, Arosio F, Mallegni C, 1990. Dose rate reduction in post-emergence weed control in sugarbeet: experimental results and technical considerations. Informatore Fitopatologico, 40(7/8):39-43.
Shalna AYu, Melamed BV, 1986. Application of bentazone-containing herbicides to barley and perennial herbage species. Khimiya v Sel'skom Khozyaistve, 24(1):44-46
Silveira EP, 1983. Integrated agricultural research on rape in RS (1980/81). Lavoura Arrozeira, 36(342):30-32, 34-36; 13 ref.
Simmonds NW, 1946. Biological flora of the British Isles. Polygonum L. em. Gaertn. Journal of Ecology, 33:117-143.
Sîrbu M, Slonovschi V, 1989. Aspects on the influence of rotations and long-term fertilizer applications on weed seed banks in the soil. Cercetari Agronomice în Moldova, 22(4):51-55; 5 ref.
Skorda EA, Adamidis T, Efthimiadis PG, 1995. Long-term effects of reduced herbicide use on weed populations and crop yield in wheat. Brighton crop protection conference: weeds. Proceedings of an international conference, Brighton, UK, 20-23 November 1995., Vol. 2:695-700; 5 ref.
Skorobogatova PA, Mirchink TG, 1985. Herbicidal activity of the microbial metabolite citrinin. Sel'skokhozyaistvennaya Biologiya, No.1:91-93; 12 ref.
Stevens OA, 1932. The number and weight of seeds produced by weeds. American Journal of Botany, 19:784-794.
Stevens OA, 1947. Studies on wild buckwheat. North Dakota Agriculture Experimental Station Bulletin, 346.
Stoimenova I, Taleva A, Mikova A, 1995. Herbicidal spectrum of some mixtures used in soybean growing. In: Proceedings, Soil Science and Strategy for Sustainable Agriculture Conference, Sofia, Bulgaria. Pochvoznanie, Agrokhimiya y Ekologyia, 30(1-6): 197-199.
Stryckers J, Himme Mvan, 1974. Comparison of several soil-applied herbicides in established Limonium Mill. plants. Stryckers, J.; Himme, M. van: Review of the results obtained for the cropping year 1972-73 by the Centrum voor Onkruidonderzoek. Rijksuniversiteit-Gent. Belgium, 152-153
Sysmans J, D'Hollander R, Schoonejans T, Tossens H, Vincinaux C, 1991. Research on herbicide efficiency and tolerance of 'low dose systems' for weed control in beet. Mededelingen van de Faculteit Landbouwwetenschappen, Rijksuniversiteit Gent, 56(3a):617-631
Talasova N, 1987. Problems with weed control in lentils. Agrochemia, 27(3):79-82
The Plant List, 2013. The Plant List: a working list of all plant species. Version 1.1. London, UK: Royal Botanic Gardens, Kew. http://www.theplantlist.org
Thomas AG, 1985. Weed survey system used in Saskatchewan for cereal and oilseed crops. Weed Science, 33(1):34-43; 6 ref.
Thomas AG, 1991. Floristic composition and relative abundance of weeds in annual crops of Manitoba. Canadian Journal of Plant Science, 71(3):831-839; 12 ref.
Thomas AG, Donaghy DI, 1991. A survey of the occurrence of seedling weeds in spring annual crops in Manitoba. Canadian Journal of Plant Science, 71(3):811-820.
Thomas AG, Wise RF, 1988. Weed survey of Manitoba cereal and oilseed crops 1986. Publication, Weed Survey Series, Agriculture Canada, No. 88-1:201pp.
Thomas AG, Wise RF, 1989. Weed surveys of Saskatchewan winter wheat fields 1985-1988. Publication, Agriculture Canada, Weed Survey Series, No. 89-1:248 pp.
Thompson DF, 1984. Observations on weed control in no-tillage crop production at Gunnedah research centre. In: Martin RJ, Felton WL, eds. No-tillage crop production in northern N.S.W. Tamworth, Australia: Department of Agriculture, 89-90.
Timson J, 1966. The germination of Polygonum convolvulus L. New Phytology, 65:423-428.
Tkachuk R, Mellish VJ, 1977. Amino acid and proximate analyses of weed seeds. Canadian Journal of Plant Science, 57(1):243-249
Toth E, Peter I, 1997. Weed control in sugarbeet with triflusulfuron-methyl based programmes: the Hungarian experience. Proceedings of the 49th international symposium on crop protection, Gent, Belgium, 6 May 1997, Part III. Mededelingen Faculteit Landbouwkundige en Toegepaste Biologische Wetenschappen, Universiteit-Gent, 1997, 62(3a): 791-798.
Townshend JL, Davidson RR, 1962. Some weed hosts of the northern root knot nematode, Meloidogyne hapla Chitwood, 1949, in Ontario. Canadian Journal of Botany, 40:543-548.
Tu HL, 1989. The evolution and control strategy of weeds in wheat fields. Proceedings, 12th Asian-Pacific Weed Science Society Conference Taipei, Taiwan: Asian-Pacific Weed Science Society, 1:67-72.
Tuganpv VV, 1973. The composition of crop and weed seeds in archeological remains of the ancient town of Osh Pando near the village of Sainino in the Mordovian ASSR (VI - IX centuries A.D.). Botanicheskii Zhurnal, 58(4):581-582
Ul'yanova TN, 1982. Kamchatka Peninsula. In: Holzner W, Numata M, eds. Biology and Ecology of weeds. The Hague, Netherlands: Dr. W. Junk, 281-284.
USDA-ARS, 1999. Germplasm Resources Information Network (GRIN). Online Database. Beltsville, Maryland, USA: National Germplasm Resources Laboratory. https://npgsweb.ars-grin.gov/gringlobal/taxon/taxonomysearch.aspx
USDA-ARS, 2015. Germplasm Resources Information Network (GRIN). Online Database. Beltsville, Maryland, USA: National Germplasm Resources Laboratory. https://npgsweb.ars-grin.gov/gringlobal/taxon/taxonomysearch.aspx
USDA-NRCS, 2015. The PLANTS Database. Baton Rouge, USA: National Plant Data Center. http://plants.usda.gov/
Valkova O, 1975. Application of some triazine compounds and propachlor. 8th International Plant Protection Congress, Reports and Informations, Section 3. Chemical control. Moscow, 2:719-720.
Veegens L, Vergracht J, 1974. Control of annual grasses and certain broad-leaved weeds in sugarbeet, maize and other field crops with Hercules 22 234. Mededelingen Fakulteit Landbouwwetenschappen, Gent, 39(2):537-553
Veselovskii IV, Saulyak PM, 1984. Herbicide mixtures based on metolachlor for maize. Khimiya v Sel'skom Khozyaistve, 12:31-32.
Wagner WL, Herbst DR, Sohmer SH, 1999. Manual of the flowering plants of Hawaii. Revised edition. Honolulu, Hawaii, USA: University of Hawaii Press/Bishop Museum Press, 1919 pp.
Webb CJ, Sykes WR, Garnock-Jones PJ, 1988. Flora of New Zealand, Volume IV: Naturalised pteridophytes, gymnosperms, dicotyledons. Christchurch, New Zealand: Botany Division, DSIR, 1365 pp.
Wells MJ, Stirton CH, 1982. South Africa. In: Holzner W, Numata M, eds. Biology and Ecology of weeds. The Hague, Netherlands: Dr. W. Junk Publishers, 339-343.
Wesolowski M, 1982. The content of weed seeds in several soils of southeast and central Poland. Part I. Bielitz soils. Annales Universitatis Marip Curie-Sklodowska, E (Agricultura), 37:9-22.
Wiesner K, 1962. Untersuchungen über den Wirtspflanzenkreis des Vergilbungsvirus der Beta-Rübe. (Corium betae Holmes). Nachrichtenblatt des Deutschen Pflanzenschutzdienstes, 16:45-53.
Willerding U, 1981. Ur- und frühgeschichtliche sowie mittelalterliche Unkrautfunde in Mitteleuropa. Zeitschrift für Pflanzenkrankheiten und Pflanzenschutz, Sonderheft, 9:65-74.
Wilson KL, 2008. Fallopia convolvulus (L.) A.Löve. New South Wales Flora Online. http://plantnet.rbgsyd.nsw.gov.au/cgi-bin/NSWfl.pl?page=nswfl&lvl=sp&name=Fallopiãconvolvulus
Wolf HW van der, 1992. The host-plants and larval cases of Coleophora therinella and C. peribenanderi (Lepidoptera: Coleophoridae). Entomologische Berichten, 52(4):47-49
Zubkov AF, 1987. Residual weed damage in winter wheat after 2,4-D application. Sovershenstvovanie khimicheskogo metoda bor'by s sornyakami Leningrad, USSR; Vsesoyuznaya Ordena Lenina i Ordena Trudovogo Krasnogo Znameni Akademiya Sel'skokhozyaistrenny kh Nauk im. V.I. Lenina, 29-34
Zuza VS, 1983. Effectiveness of herbicides applied to maize. Zashchita Rastenii, 1:27-28.
Zuza VS, 1986. On the joint action of herbicides. Agrokhimiya, No.4:79-85; 11 ref.
Zwerger P, 1987. Einfluß der Fruchtfolge, der Pflanzenbau- und Pflanzenschutz-IntensitSt auf die Populationsdynamik von UnkrSutern unter besonderer Berücksichtigung des Windenknöterichs (Fallopia convolvulus (L.) A. Löve). Dissertation UniversitSt Hohenheim. Hohenheim, Germany: University of Hohenheim.
Zwerger P, 1990. Model trials on the influence of nitrogen fertilization on the seed production and competitiveness of Fallopia convolvulus (L.) A. Love and three Polygonum-species. Zeitschrift für Pflanzenkrankheiten und Pflanzenschutz, Sonderheft, 12:107-113.
Zwerger P, 1993. Modellierung und Simulation der Populationsdynamik von annuellen UnkrSutern. Berichte aus dem Fachgebiet Herbologie der UniversitSt Hohenheim, 33.
Hassannejad, S., Ghafarbi, S. P., 2014. Weed flora survey in alfalfa (Medicago sativa L.) fields of Shabestar (northwest of Iran).Archives of Agronomy and Soil Science, 60(7) 971-991.
Macharia, I., Backhouse, D., Wu, S. B., Ateka, E. M., 2016. Weed species in tomato production and their role as alternate hosts of Tomato spotted wilt virus and its vector Frankliniella occidentalis.Annals of Applied Biology, 169(2) 224-235.
Oh, S. M., Moon, B. C., Kim, C. S., 2007. Current status on influx and habitat of exotic weeds in Korea. In: Proceedings of the 21st Asian Pacific Weed Science Society (APWSS) Conference, 2-6 October 2007, Colombo, Sri Lanka [Proceedings of the 21st Asian Pacific Weed Science Society (APWSS) Conference, 2-6 October 2007, Colombo, Sri Lanka.], [ed. by Marambe, B, Sangakkara, U. R., Costa, W. A. J. M. de, Abeysekara, A. S. K.]. Peradeniya, Sri Lanka: Asian Pacific Weed Science Society. 608-613.
Moskova, T., Dimitrov, G., Tityanov, M., 2018. Distribution and degree of weed growth of amaranth and other weeds in sunflower crops in Plovdiv and Stara Zagora regions.Journal of Mountain Agriculture on the Balkans, 21(1) 158-168. http://www.rimsa.eu/images/forage_production_vol_21-1_part_2_2018.pdf
Woźniak, A., 2020. Effect of cereal monoculture and tillage systems on grain yield and weed infestation of winter durum wheat.International Journal of Plant Production, 14(1) 1-8.
Mohammadi, A. H., Haghdel, M., Mohammadi-Moghaddam, M., Banihashemi, Z., 2006. Current status of Verticillium wilt disease of pistachio trees in Iran.Acta Horticulturae, No.726631-635. http://www.actahort.org
Goerke, K., Schönhammer, A., Schulte, M., Gerowitt, B., 2007. Weeds in oilseed rape in Germany - status and assessment of changes. In: European Weed Research Society, 14th EWRS Symposium, Hamar, Norway, 17-21 June 2007 [European Weed Research Society, 14th EWRS Symposium, Hamar, Norway, 17-21 June 2007.], [ed. by Fløistad, E.]. Doorwerth, Netherlands: European Weed Research Society. 198. http://www.ewrs-symposium2007.com
Stobbs, L. W., Greig, N., Weaver, S., Shipp, L., Ferguson, G., 2009. The potential role of native weed species and bumble bees (Bombus impatiens) on the epidemiology of Pepino mosaic virus.Canadian Journal of Plant Pathology, 31(2) 254-261. http://www.tandfonline.com/doi/abs/10.1080/07060660909507599
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