Abbott I, 1993. Insect pest problems of eucalypt plantations in Australia. 6. Western Australia. Australian Forestry, 56(4):381-384; 21 ref.

Ball OJP, Miles CO, Prestidge RA, 1997. Ergopeptine alkaloids and Neotyphodium lolii-mediated resistance in perennial ryegrass against adult Heteronychus arator (Coleoptera: Scarabaeidae). Journal of Economic Entomology, 90(5):1382-1391; 39 ref.

Blank RH, Olson MH, Bell DS, 1982. Insecticidal control of black beetle (Heteronychus arator) in pasture. New Zealand Journal of Experimental Agriculture, 10(4):433-438

CABI/EPPO, 2000. Heteronychus arator. Distribution Maps of Plant Pests, Map No. 163. Wallingford, UK: CAB International.

Cumpston DM, 1940. On the external morphology and biology of Heteronychus sanctae-Helenae Blanch. And Metanastes vulgivagus Olliff (Col., Scarabaeidae, Dynastinae). Proceedings of the Linnaean Society of New South Wales, 65:289-300.

Drinkwater TW, 1979. The black maize beetle. Farming in South Africa Maize Series D: Leaflet D4/1979.

Drinkwater TW, 1982. The control of the black maize beetle, Heteronychus arator (Col.: Scarabpidae), in maize in South Africa. Phytophylactica, 14(4):165-167

Drinkwater TW, 1987. The efficacy of insecticides in the control of the black maize beetle, Heteronychus arator (Col.: Scarabpidae) in maize in South Africa. Phytophylactica, 19(3):275-277

Drinkwater TW, Groenewald LH, 1994. Comparison of imidacloprid and furathiocarb seed dressing insecticides for the control of the black maize beetle, Heteronychus arator Fabricius (Coleoptera: Scarabaeidae), in maize. Crop Protection, 13(6):421-424

EPPO, 2014. PQR database. Paris, France: European and Mediterranean Plant Protection Organization. http://www.eppo.int/DATABASES/pqr/pqr.htm

Endrödi S, 1985. The Dynastinae of the World. Dordrecht, Netherlands: Dr. W. Junk, 800 pp.

Jenkins CFH, 1965. The black beetle. Journal of Agriculture Western Australia, 6:39-42.

King PD, 1977. Effect of plant species and organic matter on feeding behaviour and weight gain of larval black beetle, Heteronychus arator (Coleoptera: Scarabpidae). New Zealand Journal of Zoology, 4(4):445-448

King PD, Meekings JS, Mercer CF, 1982. Effects of whitefringed weevil (Graphognathus leucoloma) and black beetle (Heteronychus arator) populations on pasture species. New Zealand Journal of Agricultural Research, 25(3):405-414

King PD, Mercer CF, Meekings JS, 1981. Ecology of black beetle, Heteronychus arator (Coleoptera: Scarabpidae) - influence of pasture species on oviposition site preference. New Zealand Journal of Zoology, 8(1):119-122

King PD, Mercer CF, Meekings JS, 1981. Ecology of black beetle, Heteronychus arator (Coleoptera: Scarabpidae) - population modelling. New Zealand Journal of Agricultural Research, 24(1):99-105

King PD, Mercer CF, Meekings JS, 1981. Ecology of black beetle, Heteronychus arator (Coleoptera: Scarabpidae) - population studies. New Zealand Journal of Agricultural Research, 24(1):87-97

Landin BO, 1964. The identity of 'Scarabaeus arator Fabricius, 1775' (Col. Lamellicornia) with the designation of neotypes of arator Illig. and a list of the insects from Cape Colony described by Fabricius 1775. Opuscula Entomology, 29:117-142.

Learmonth SE, Sproul AN, 1988. Pasture rotation - the main source of insect pest problems in Western Australian potato crops. In: Stahle PP, ed. Proceedings of the 5th Australasian Conference on Grassland Invertebrate Ecology. Melbourne, Australia: D & D Printing, 163-168.

Matthiessen JN, 1993. African black beetle population regulation in a Mediterranean climate region of Australia - an enigma. In: Prestidge RA, ed. Proceedings of the 6th Australasian Conference on Grassland Invertebrate Ecology. Hamilton, New Zealand: AgResearch, 60-66.

Matthiessen JN, Learmonth SE, 1995. Impact of the soil insects African black beetle, Heteronychus arator (Coleoptera: Scarabaeidae) and whitefringed weevil, Graphognathus leucoloma (Coleoptera: Curculionidae), on potatoes and effects of soil insecticide treatments in south-western Australia. Bulletin of Entomological Research, 85(1):101-111

Matthiessen JN, Learmonth SE, 1998. Seasonally contrasting activity of African black beetle, Heteronychus arator (Coleoptera: Scarabaeidae): implications for populations, pest status and management. Bulletin of Entomological Research, 88(4):443-450; 18 ref.

Matthiessen JN, Learmonth SE, Sumner MW, 1997. Surface activity of African black beetle in spring favours a pre-oviposition adulticide approach to its management. In: Allsopp PG, Rogers DJ, Robertson LN, eds. Soil invertebrates in 1997. Proceedings of the 3rd Brisbane Workshop on Soil Invertebrates. Indooroopilly, Australia: Bureau of Sugar Experiment Stations, 149-152.

Matthiessen JN, Ridsdill-Smith TJ, 1991. Populations of African black beetle, Heteronychus arator (Coleoptera: Scarabpidae) in a Mediterranean climate region of Australia. Bulletin of Entomological Research, 81(1):85-91

Russell GB, Sutherland ORW, Christmas PE, Wright H, 1982. Feeding deterrents for black beetle larvae, Heteronychus arator (Scarabpidae), in Trifolium repens. New Zealand Journal of Zoology, 9(1):145-149

Sainty BJ, 1991. Experiences with African black beetle and weevils. Australian & New Zealand Wine Industry Journal, 6(3):237-238

Sutherland ORW, 1976. A chemical basis for plant resistance to grass grub and black beetle larvae (Coleoptera: Scarabpidae). Proceedings of the New Zealand Grassland Association, 37(1):126-131

Sutherland ORW, Greenfield WJ, 1978. Effect of root extracts of resistant pasture plants on the feeding and survival of black beetle larvae, Heteronychus arator (Scarabpidae). New Zealand Journal of Zoology, 5(1):173-175

Taylor TA, 1951. The black maize beetle (Heteronychus sanctae-helenae). South African Farming, 26:299-302.

Toit HA Du, 1997. Habitat preference of African black beetle and other soil insect pests of maize in South Africa. In: Allsopp PG, Rogers DJ, Robertson LN, eds. Soil invertebrates in 1997. Proceedings of the 3rd Brisbane Workshop on Soil Invertebrates. Indooroopilly, Australia: Bureau of Sugar Experiment Stations, 44-47.

Watson RN, 1980. Dispersal and distribution of Heteronychus arator in New Zealand (Coleoptera: Scarabpidae). In: Crosby TK, Pottinger RP, ed. Proceedings of the 2nd Australasian conference on grassland invertebrate ecology. Palmerston North, New Zealand 22-26 May 1978. Government Printer. Wellington New Zealand, 149-152

Wright WE, 1958. The black beetle. The Agricultural Gazette, August:422-427.

Bulinski, J., Matthiessen, J. N., Alexander, R., 2006. Development of a cost-effective, pesticide-free approach to managing African black beetle (Heteronychus arator) in Australian eucalyptus plantations.Crop Protection, 25(11) 1161-1166.

Hume, D. E., Ryan, D. L., Cooper, B. M., Popay, A. J., 2007. Agronomic performance of AR37-infected ryegrass in northern New Zealand.Proceedings of the New Zealand Grassland Association, 69201-205.

Matthiessen, J. N., Learmonth, S. E., 1993. Spatial sampling of insects, plant parts and insect attacks in the soil of potato crops.Bulletin of Entomological Research, 83(4) 607-612.

Venter, R. J. H., Louw, M., 1978. Heteronychus arator (Fabricius), a potentially dangerous pest of potatoes (Coleoptera: Scarabaeidae).Phytophylactica, 10(3) 99.