Capra hircus (goats)
Datasheet Types: Natural enemy, Livestock species, Invasive species, Host animal
Abstract
This datasheet on Capra hircus covers Identity, Overview, Distribution, Dispersal, Biology & Ecology, Physiology, Impacts, Uses, Prevention/Control, Management, Further Information.
Identity
- Preferred Scientific Name
- Capra hircus Linnaeus 1758
- Preferred Common Name
- goats
- Other Scientific Names
- Capra aegagrus hircus
- Capra indicus
- International Common Names
- Englishbillygoatkidkidsnanny
- Spanishcabras
- Frenchchèvres
- Local Common Names
- GermanyHausziegeZiegen
- Italycapre
- Portugalcabras
Pictures

Goats grazing in a tree
Capra hircus (goats); goats grazing in an Argan tree (Argania spinosa). Essaouira, Morocco. June 2013.
©Grand Parc - Bordeaux, France/via wikipedia - CC BY 2.0

Herd of dairy goats
Capra hircus (goats); herd of Saanen dairy goats. Lukoz Farm, Russia. September 2008.
©Efras/via wikipedia - CC BY-SA 3.0

Herd of Tibetan goats
Capra hircus (goats); herd of Tibetan goats, with their goatherd. Tibet. October 2005.
©Adam Schorsch/via wikipedia - CC BY 2.0

Female
Capra hircus (goats); female Toggenburg goat from "Zlatusha" farm of the Institute of Animal Science, Kostinbrod, Bulgaria, 1995.
©Nadezhda Petrova
Summary of Invasiveness
The goat ( Capra hircus ) was domesticated 10,000 years ago in the highlands of western Iran. These herbivores have a highly varied diet and are able to utilise a larger number of plant species than other livestock. Goats alter plant communities and forest structure and threaten vulnerable plant species. The reduction of vegetation reduces shelter options for native animals and overgrazing in native communities leads to ecosystem degradation. Feral goats spread disease to native animals. Native fauna on islands are particularly susceptible.
Taxonomic Tree
Breeds
Description
Males weigh between 45 and 55 kilos and females weigh between 25 and 35 kilos. Colouration is highly variable from mostly black, to various shades of brown, and from single-coloured to multi-coloured. Black anterior with brown posterior is a common pattern. Horns are dimorphic, having homonymous spiral and anterior keel. Males are bearded and produce pheromones during the breeding season.
Pathogens Carried
Distribution
Goats were domesticated 10,000 years ago in the highlands of western Iran. Feral goats are derived from domestic goats, originally native to Asia.
Distribution Map
Distribution Table
Means of Movement and Dispersal
Introduction pathways to new locations
Other: Goats ( Capra hircus ) were released as potential food for people marooned by shipwrecks.
Transportation of domesticated animals: Goats ( Capra hircus ) were introduced to islands for their milk.
Local dispersal methods
Escape from confinement:
Natural dispersal (local):
Other: Goats ( Capra hircus ) were released as potential food for people marooned by shipwrecks.
Transportation of domesticated animals: Goats ( Capra hircus ) were introduced to islands for their milk.
Local dispersal methods
Escape from confinement:
Natural dispersal (local):
Pathway Causes
Pathway cause | Notes | Long distance | Local | References |
---|---|---|---|---|
Animal production (pathway cause) | Yes | |||
Escape from confinement or garden escape (pathway cause) | Yes | |||
Food (pathway cause) | Yes | |||
Self-propelled (pathway cause) | Yes |
Habitat
Goats usually move in herds that roam over territories up to 20km wide. Sometimes herd ranges can be as small as 100m 2 . Males usually wander more widely than females. Grasslands, scrub lands, rocky outcrops and semi-open or open forests are all used extensively by goats as habitat substrate. In bad weather, they may seek shelter under rocky ledges (SPREP, 2000).
Habitat List
Category | Sub category | Habitat | Presence | Status |
---|---|---|---|---|
Terrestrial | Terrestrial – Managed | Managed forests, plantations and orchards | Present, no further details | Harmful (pest or invasive) |
Terrestrial | Terrestrial ‑ Natural / Semi-natural | Natural forests | Present, no further details | Harmful (pest or invasive) |
Terrestrial | Terrestrial ‑ Natural / Semi-natural | Natural grasslands | Present, no further details | Harmful (pest or invasive) |
Terrestrial | Terrestrial ‑ Natural / Semi-natural | Scrub / shrublands | Present, no further details | Harmful (pest or invasive) |
Biology and Ecology
Nutrition
Goats are herbivores and will forage on any palatable plants in their home range. Goats have rather large rumino-reticular volume so they are able to subsist on poorer quality plants than most herbivores, therefore goats can survive and subsist in heavily exploited environments.
Goats are herbivores and will forage on any palatable plants in their home range. Goats have rather large rumino-reticular volume so they are able to subsist on poorer quality plants than most herbivores, therefore goats can survive and subsist in heavily exploited environments.
In a study conducted by Chimera, Coleman and Parkes (1995) the rumen contents of 49 goats (captured in 1989) were identified and the dry weight of each component was measured to produce a breakdown of the diet of a small, unique remnant population (now extinct) of feral goats on Auckland Island (a Sub-Antarctic island located south of New Zealand). Woody plants and grasses made up the bulk of their diet (41% and 39%, respectively); seaweeds made up 13%, ferns 4% and herbaceous species 3%. At least 40 plant species were eaten by the goats, but just three species made up half of the total: rata ( Metrosideros umbellate ), snow tussock ( Chionochloa Antarctica ) and kelp ( Durvillea Antarctica ). Rata ( Metrosideros umbellate ), Coprosma foetidissima, Pseudopanax simplex and Carex appressa were the most commonly found species - eaten in at least some quantity by 90%, 80%, 76% and 69% of the goats respectively.
Unidentified grasses composed 21.4% of the total contents (dry weight) and were found in 100% of goats. NB: This population had a unique genetic makeup as it evolved separately for over 100 years on an isolated island with a cold and harsh climate and so it may not represent the dietary preferences of all goats. However, it highlights the huge adaptability of goats and their ability to subsist in particular and inhospitable landscapes.
Reproduction
Both sexes are physiologically capable of reproduction at about 6 months of age. Dominant males fight to win females. They follow a serial pattern and attend to one female after another as they come into oestrus. In one sense this is termed polygyny, as males breed with as many females as they can during a breeding period, but more properly this is serial monogamy as a male will tend a female for extended periods, both before and after copulation and before leaving in search of a different mate.
Both sexes are physiologically capable of reproduction at about 6 months of age. Dominant males fight to win females. They follow a serial pattern and attend to one female after another as they come into oestrus. In one sense this is termed polygyny, as males breed with as many females as they can during a breeding period, but more properly this is serial monogamy as a male will tend a female for extended periods, both before and after copulation and before leaving in search of a different mate.
Realised reproduction varies among populations, seasons and years. At best a typical female goat would produce one young in its first pregnancy and twin kids in subsequent pregnancies. However, most females are at less than optimal condition, which may result in a twinning rate of as little as 0%. At best, herds may produce twins at a rate of 80% or more. Triplets are not uncommon.
Physiology Table
Type | Value |
Heart rate (beats per minute) | 70 - 135 |
Daily urine volume (mL/kg of body weight) | 10 - 40 |
Rectal temp (?C+0.5?C) | 39 |
Haematology
Type | Value |
---|---|
Haemoglobin (g/dL) | 8 - 12 |
PCV (%) | 22 - 38 |
RBC (x10 6 /µL) | 8 - 18 |
MCV (fL) | 16 - 25 |
MCH (pg) | 5 - 8 |
MCHC (g/dL) | 30 - 36 |
Reticulocytes (% of RBC) | 0 |
WBC (/µL) | 4000 - 13000 |
Neutrophils (mature) (/µL) | 1200 - 7200 |
Lymphocytes (/µL) | 2000 - 9000 |
Monocytes (/µL) | 0 - 550 |
Eosinophils (/µL) | 50 - 650 |
Basophils (/µL) | 0 - 120 |
Platelets (x10 5 /µL) | 3 - 6 |
Plasma proteins (g/dL) | 6 - 7.5 |
Fibrinogen (mg/dL) | 100 - 400 |
Blood Chemistry
Type | Value |
---|---|
Alkaline phosphatase (U/L) | 42 - 775 |
Aspartate aminotranferase (U/L) | 67 - 117 |
Bicarbonate (mmol/L) | 26 - 30 |
Bilirubin - total (mg/dL) | 0 - 0.1 |
Calcium (mg/dL) | 8.5 - 10.2 |
Chloride (mEq/L) | 99 - 110 |
Cholesterol (mg/dL) | 80 - 130 |
γ-glutamyltransferase (U/L) | 43 - 71 |
Glucose (mg/dL) | 58 - 76 |
Magnesium (mg/dL) | 2.8 - 3.6 |
Potassium (mEq/L) | 3.5 - 6.7 |
Sodium (mEq/L) | 142 - 155 |
Sorbitol dehydrogenase (U/L) | 35 - 80 |
Urea Nitrogen (mg/dL) | 15 - 33 |
Total Protein (g/dL) | 5.9 - 7.4 |
Albumin (g/dL) | 2.7 - 3.9 |
a1 globulin (g/dL) | 0.5 - 0.7 |
β1 globulin (g/dL) | 0.7 - 1.2 |
β2 globulin (g/dL) | 0.3 - 0.6 |
γ1 globulin (g/dL) | 0.9 - 3 |
List of Diseases and Disorders
Natural enemy of
Impact Summary
Category | Impact |
---|---|
Biodiversity (generally) | Negative |
Environment (generally) | Negative |
Livestock production | Negative |
Impact
General Impacts
Original text compiled by IUCN SSC Invasive Species Specialist Group (ISSG)
Biodiversity on islands is vulnerable to introduced species, and the vegetation tends to lack adaptions to mammalian herbivores (Campbell and Donlan 2005). Unfortunately, goats (
Capra hircus
) have been established on many islands. Goats alter plant communities and forest structures and threaten vulnerable plant species; the flow-down effect of these outcomes includes increased soil erosion and the reduction of native fauna that share a similar environmental niche (Spatz and Mueller-Dombois 1973, Coblentz 1978, Parkes 1984, Brennan 1986, Coblentz and Van Vuren 1987, Cronk 1989, Walker 1991, Moran 1996, Desender
et al
. 1999, in Campbell and Donlan 2005). In some island ecosystems it has been the case that goats are the most destructive herbivore present (King, 1985). Feral goats are particularly destructive in such environments and cause a huge loss of native vegetation due to their grazing habits. This leads to ecosystem degradation and biodiversity loss (Coblentz 1978; Schofield 1989; Moran 1996; Desender
et al.
1999 in Campbell and Donlan 2005).
Threatened Species
Threatened species | Where threatened | Mechanisms | References | Notes |
---|---|---|---|---|
Acrocephalus luscinia (nightingale reed warbler) | Guam Northern Mariana Islands | Herbivory/grazing/browsing | ||
Adelocosa anops (Kauai cave wolf spider) | Hawaii | Herbivory/grazing/browsing | ||
Bidens amplectens (koko'olau) | Hawaii | Herbivory/grazing/browsing | ||
Bidens campylotheca subsp. waihoiensis (ko`oko`olau) | Hawaii | Herbivory/grazing/browsing | ||
Bidens conjuncta (ko`oko`olau) | Hawaii | Herbivory/grazing/browsing | ||
Bidens micrantha subsp. ctenophylla (ko`oko`olau) | Hawaii | Herbivory/grazing/browsing | ||
Calamagrostis hillebrandii (Hillegrands reedgrass) | Hawaii | Herbivory/grazing/browsing | ||
Canavalia pubescens (jack bean) | Hawaii | Herbivory/grazing/browsing | ||
Cinclodes aricomae | Peru | Herbivory/grazing/browsing | ||
Cyanea asplenifolia (haha) | Hawaii | Herbivory/grazing/browsing | ||
Cyanea calycina | Hawaii | Herbivory/grazing/browsing | ||
Cyanea duvalliorum | Hawaii | Herbivory/grazing/browsing | ||
Cyanea horrida (prickly cyanea) | Hawaii | Herbivory/grazing/browsing | ||
Cyanea kunthiana (Kunth's cyanea) | Hawaii | Herbivory/grazing/browsing | ||
Cyanea lanceolata (lanceleaf cyanea) | Hawaii | Herbivory/grazing/browsing | ||
Cyanea magnicalyx | Hawaii | Herbivory/grazing/browsing | ||
Cyanea maritae | Hawaii | Herbivory/grazing/browsing | ||
Cyanea marksii (Marks' cyanea) | Hawaii | Herbivory/grazing/browsing | ||
Cyanea mauiensis | Hawaii | Herbivory/grazing/browsing | ||
Cyanea munroi | Hawaii | Herbivory/grazing/browsing | ||
Cyanea obtusa (bluntlobe cyanea) | Hawaii | Herbivory/grazing/browsing | ||
Cyanea profuga (Mapulehu Valley cyanea) | Hawaii | Herbivory/grazing/browsing | ||
Cyanea recta (Kealia cyanea) | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Cyanea solanacea (popolo) | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Cyclura stejnegeri (Mona Island iguana) | Puerto Rico | Competition - monopolizing resources | ||
Cyrtandra ferripilosa | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Cyrtandra filipes | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Cyrtandra limahuliensis (Limahuli cyrtandra) | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Cyrtandra oxybapha | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Doryopteris takeuchii (Takeuch's lipfern) | Hawaii | Herbivory/grazing/browsing | ||
Dracaena forbesii (hala pepe) | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Chrysodracon hawaiiensis (hala pepe) | Hawaii | Herbivory/grazing/browsing | ||
Drosophila aglaia | Hawaii | Herbivory/grazing/browsing | ||
Drosophila differens (Hawaiian picture-wing fly) | Hawaii | Herbivory/grazing/browsing | ||
Drosophila digressa | Hawaii | Herbivory/grazing/browsing | ||
Drosophila hemipeza | Hawaii | Herbivory/grazing/browsing | ||
Drosophila heteroneura | Hawaii | Herbivory/grazing/browsing | ||
Drosophila montgomeryi | Hawaii | Herbivory/grazing/browsing | ||
Drosophila musaphilia | Hawaii | Herbivory/grazing/browsing | ||
Drosophila ochrobasis | Hawaii | Herbivory/grazing/browsing | ||
Drosophila substenoptera | Hawaii | Herbivory/grazing/browsing | ||
Emballonura semicaudata rotensis (Pacific sheath-tailed bat) | Northern Mariana Islands | Ecosystem change / habitat alteration | ||
Epicrates monensis monensis (Mona boa) | Puerto Rico | Ecosystem change / habitat alteration | ||
Festuca hawaiiensis (Hawai'i fescue) | Hawaii | Herbivory/grazing/browsing Ecosystem change / habitat alteration | ||
Festuca molokaiensis | Hawaii | Herbivory/grazing/browsing Ecosystem change / habitat alteration | ||
Gardenia remyi (Remy's gardenia) | Hawaii | Herbivory/grazing/browsing Ecosystem change / habitat alteration | ||
Harrisia portoricensis (higo chumbo) | Puerto Rico | Herbivory/grazing/browsing | ||
Huperzia stemmermanniae (Stemmermann's clubmoss) | Hawaii | Herbivory/grazing/browsing Ecosystem change / habitat alteration | ||
Hylaeus assimulans (assimulans yellow-faced bee) | Hawaii | Herbivory/grazing/browsing Ecosystem change / habitat alteration | ||
Hylaeus facilis (easy yellow-faced bee) | Hawaii | Herbivory/grazing/browsing Ecosystem change / habitat alteration | ||
Hylaeus hilaris (hilaris yellow-faced bee) | Hawaii | Herbivory/grazing/browsing Ecosystem change / habitat alteration | ||
Hylaeus kuakea (Hawaiian yellow-faced bee) | Hawaii | Herbivory/grazing/browsing Ecosystem change / habitat alteration | ||
Hylaeus longiceps | Hawaii | Herbivory/grazing/browsing Ecosystem change / habitat alteration | ||
Hylaeus mana | Hawaii | Herbivory/grazing/browsing Ecosystem change / habitat alteration | ||
Isodendrion longifolium (longleaf isodendrion) | Hawaii | Herbivory/grazing/browsing Ecosystem change / habitat alteration | ||
Joinvillea ascendens ascendens | Hawaii | Herbivory/grazing/browsing Ecosystem change / habitat alteration | ||
Korthalsella taenioides (Degener's korthal mistletoe) | Hawaii | Herbivory/grazing/browsing Ecosystem change / habitat alteration | ||
Leiolopisma telfairii (Round Island skink) | Mauritius | Ecosystem change / habitat alteration | ||
Lobelia monostachya (Waianae Range lobelia) | Hawaii | Herbivory/grazing/browsing | ||
Lobelia niihauensis (Niihau lobelia) | Hawaii | Herbivory/grazing/browsing | ||
Loxioides bailleui (palila) | Hawaii | Herbivory/grazing/browsing | ||
Melanthera tenuifolia (Waianae Range nehe) | Hawaii | Herbivory/grazing/browsing | ||
Melanthera venosa (spreading nehe) | Hawaii | Herbivory/grazing/browsing | ||
Melanthera waimeaensis (Waimea Canyon nehe) | Hawaii | Herbivory/grazing/browsing | ||
Melicope christophersenii | Hawaii | Herbivory/grazing/browsing Ecosystem change / habitat alteration | ||
Melicope hiiakae | Hawaii | Herbivory/grazing/browsing Ecosystem change / habitat alteration | ||
Melicope makahae | Hawaii | Ecosystem change / habitat alteration | ||
Melicope makahae | Hawaii | Ecosystem change / habitat alteration | ||
Mucuna sloanei var. persericea | Hawaii | Herbivory/grazing/browsing Ecosystem change / habitat alteration | ||
Myrsine linearifolia (narrowleaf colicwood) | Hawaii | Competition - monopolizing resources | ||
Nothocestrum latifolium (broadleaf aiea) | Hawaii | Herbivory/grazing/browsing Ecosystem change / habitat alteration | ||
Nothocestrum peltatum (Oahu aiea) | Hawaii | Herbivory/grazing/browsing | ||
Nototrichium humile (kaala rockwort) | Hawaii | Herbivory/grazing/browsing | ||
Oceanodroma castro (band-rumped storm-petrel) | Montana | Ecosystem change / habitat alteration | ||
Ochrosia haleakalae (island yellowwood) | Hawaii | Herbivory/grazing/browsing Ecosystem change / habitat alteration | ||
Ochrosia kilaueaensis (Hawai'i yellowwood) | Hawaii | Herbivory/grazing/browsing | ||
Oreomystis bairdi (akikiki) | Hawaii | Herbivory/grazing/browsing | ||
Oreomystis mana (Hawaii creeper) | Hawaii | Herbivory/grazing/browsing | ||
Osmoxylon mariannense | Northern Mariana Islands | Herbivory/grazing/browsing | ||
Ovis ammon (argali) | Kyrgyzstan Mongolia Tajikistan | Competition - monopolizing resources | ||
Ovis canadensis sierrae (Sierra Nevada bighorn sheep) | California | Pathogenic | ||
Panicum fauriei (Carter's panicgrass) | Hawaii | Ecosystem change / habitat alteration | ||
Paroreomyza flammea (Molokai creeper) | Hawaii | Ecosystem change / habitat alteration | ||
Partula gibba (humped tree snail) | Guam Northern Mariana Islands | Ecosystem change / habitat alteration | ||
Partula langfordi (Langford's tree snail) | Northern Mariana Islands | Ecosystem change / habitat alteration | ||
Partula radiolata (Guam tree snail) | Guam | Ecosystem change / habitat alteration | ||
Partulina semicarinata (Lanai tree snail) | Hawaii | Ecosystem change / habitat alteration | ||
Partulina variabilis (Lanai tree snail) | Hawaii | Ecosystem change / habitat alteration | ||
Peucedanum sandwicense (makou) | Hawaii | Herbivory/grazing/browsing | ||
Phyllostegia glabra var. lanaiensis (ulihi phyllostegia) | Hawaii | Ecosystem change / habitat alteration | ||
Phyllostegia knudsenii (Waimea phyllostegia) | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Phyllostegia mannii (Mann's phyllostegia) | Hawaii | Ecosystem change / habitat alteration | ||
Phyllostegia mollis (Waianae Range phyllostegia) | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Phyllostegia parviflora (smallflower phyllostegia) | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Phyllostegia renovans (red-leaf phyllostegia) | Hawaii | Herbivory/grazing/browsing | ||
Phyllostegia waimeae (Kauai phyllostegia) | Hawaii | Herbivory/grazing/browsing | ||
Phyllostegia wawrana (fuzzystem phyllostegia) | Hawaii | Herbivory/grazing/browsing | ||
Phytotoma raimondii (Peruvian plantcutter) | Peru | Ecosystem change / habitat alteration | ||
Pittosporum napaliense (royal cheesewood) | Hawaii | Herbivory/grazing/browsing | ||
Plantago hawaiensis (Hawai'i plantain) | Hawaii | Herbivory/grazing/browsing | ||
Plantago princeps | Hawaii | Herbivory/grazing/browsing | ||
Platydesma cornuta var. cornuta | Hawaii | Herbivory/grazing/browsing Ecosystem change / habitat alteration | ||
Platydesma cornuta var. decurrens | Hawaii | Herbivory/grazing/browsing Ecosystem change / habitat alteration | ||
Platydesma rostrata | Hawaii | Herbivory/grazing/browsing Ecosystem change / habitat alteration | ||
Poa mannii (Mann's bluegrass) | Hawaii | Herbivory/grazing/browsing | ||
Portulaca sclerocarpa | Hawaii | Herbivory/grazing/browsing | ||
Pritchardia hardyi (Makaleha pritchardia) | Hawaii | Herbivory/grazing/browsing | ||
Pritchardia kaalae (Waianae Range pritchardia) | Hawaii | Herbivory/grazing/browsing | ||
Pritchardia munroi (Kamalo pritchardia) | Hawaii | Herbivory/grazing/browsing | ||
Pritchardia napaliensis | Hawaii | Herbivory/grazing/browsing | ||
Pritchardia remota (Remota loula palm) | Hawaii | Herbivory/grazing/browsing | ||
Pseudognaphalium sandwicensium var. molokaiense (enaena) | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Psittirostra psittacea (Ou) | Hawaii | Herbivory/grazing/browsing | ||
Psychotria grandiflora (large-flowered balsamo) | Hawaii | Herbivory/grazing/browsing | ||
Psychotria hobdyi (Hobdy's wild-coffee) | Hawaii | Herbivory/grazing/browsing | ||
Pteralyxia kauaiensis (Kauai pteralyxia) | Hawaii | Herbivory/grazing/browsing | ||
Pteralyxia laurifolia (ridged pteralyxia) | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Pterodroma sandwichensis | Hawaii | Herbivory/grazing/browsing | ||
Pteropus mariannus mariannus (Mariana fruit bat) | Guam Northern Mariana Islands | Ecosystem change / habitat alteration | ||
Ranunculus hawaiensis (Hawai'ian buttercup) | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Ranunculus mauiensis (Maui buttercup) | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Remya kauaiensis (Kauai remya) | Hawaii | Ecosystem change / habitat alteration | ||
Remya mauiensis (Maui remya) | Hawaii | Competition (unspecified) | ||
Remya montgomeryi (Kalalau Valley remya) | Hawaii | Ecosystem change / habitat alteration | ||
Sanicula mariversa (Waianae Range blacksnakeroot) | Hawaii | Ecosystem change / habitat alteration | ||
Schiedea apokremnos (Kauai schiedea) | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Schiedea haleakalensis (Haleakala schiedea) | Hawaii | Ecosystem change / habitat alteration | ||
Schiedea helleri (Kaholuamanu schiedea) | Hawaii | Ecosystem change / habitat alteration | ||
Schiedea helleri (Kaholuamanu schiedea) | Hawaii | Ecosystem change / habitat alteration | ||
Schiedea hookeri (sprawling schiedea) | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Schiedea jacobii (Jacobi's schiedea) | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Schiedea kaalae (Oahu schiedea) | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Schiedea kauaiensis (Kauai schiedea) | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Schiedea kealiae (Waianae Range schiedea) | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Schiedea lydgatei (Kamalo Gulch schiedea) | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Schiedea membranacea | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Schiedea nuttallii | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Schiedea pubescens | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Schiedea salicaria | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Schiedea sarmentosa | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Schiedea spergulina var. leiopoda | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Schiedea spergulina var. spergulina | Hawaii | Herbivory/grazing/browsing | ||
Schiedea stellarioides | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Sesbania tomentosa | Hawaii | Ecosystem change / habitat alteration | ||
Sibara filifolia (Santra Cruz Island Rockcress) | California | Herbivory/grazing/browsing | ||
Silene alexandri | Hawaii | Ecosystem change / habitat alteration | ||
Silene hawaiiensis (Hawaii catchfly) | Hawaii | Herbivory/grazing/browsing | ||
Silene lanceolata (Kauai catchfly) | Hawaii | Ecosystem change / habitat alteration | ||
Solanum incompletum | Hawaii | Ecosystem change / habitat alteration | ||
Solanum sandwicense | Hawaii | Ecosystem change / habitat alteration | ||
Spermolepis hawaiiensis (Hawaii scaleseed) | Hawaii | Ecosystem change / habitat alteration | ||
Stenogyne campanulata (Kalalau Valley stenogyne) | Hawaii | Herbivory/grazing/browsing | ||
Stenogyne kanehoana (Oahu stenogyne) | Hawaii | Herbivory/grazing/browsing | ||
Stenogyne purpurea (purplefruit stenogyne) | Hawaii | Herbivory/grazing/browsing | ||
Streptanthus bracteatus (bracted twistflower) | Texas | Herbivory/grazing/browsing | ||
Synthliboramphus hypoleucus (Xantus' murrelet) | Canada Mexico California Oregon | Ecosystem change / habitat alteration | ||
Tetramolopium arenarium (Maui tetramolopium) | Hawaii | Herbivory/grazing/browsing | ||
Tetramolopium filiforme (ridgetop tetramolopium) | Hawaii | Herbivory/grazing/browsing | ||
Tetramolopium lepidotum (Waianae Range tetramolopium) | Hawaii | Herbivory/grazing/browsing | ||
Tetramolopium remyi (Awalua Ridge tetramolopium) | Hawaii | Herbivory/grazing/browsing | ||
Tetraplasandra flynnii | Hawaii | Herbivory/grazing/browsing | ||
Urera kaalae | Hawaii | Ecosystem change / habitat alteration Pest and disease transmission | ||
Urocyon littoralis catalinae (Santa Catalina Island fox) | California | Ecosystem change / habitat alteration | ||
Verbesina dissita (big-leaved crownbeard) | California | Herbivory/grazing/browsing | ||
Vetericaris chaceorum (anchialine pool shrimp) | Hawaii | Ecosystem change / habitat alteration | ||
Vicia menziesii (Hawaiian vetch) | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Vigna o-wahuensis (Oahu cowpea) | Hawaii | Herbivory/grazing/browsing | ||
Viola chamissoniana subsp. chamissoniana (pamakani) | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Wilkesia hobdyi (dwarf iliau) | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing | ||
Zanthoxylum dipetalum var. tomentosum | Hawaii | Ecosystem change / habitat alteration Herbivory/grazing/browsing |
Risk and Impact Factors
Invasiveness
Proved invasive outside its native range
Highly adaptable to different environments
Capable of securing and ingesting a wide range of food
Impact outcomes
Ecosystem change/ habitat alteration
Negatively impacts agriculture
Negatively impacts forestry
Negatively impacts animal health
Negatively impacts livelihoods
Reduced native biodiversity
Threat to/ loss of native species
Impact mechanisms
Competition - monopolizing resources
Competition (unspecified)
Pest and disease transmission
Herbivory/grazing/browsing
Interaction with other invasive species
Pathogenic
Uses
The ability of goat herds to survive in harsh environments has led to their introduction onto many islands, including Saint Helena Island, the Juan Fernandez Islands and Hawaii. They provided food for colonising people (specifically European colonisers and ship crews) (Campbell and Donlan 2005). Fishermen may have spread goats onto new islands, such as San Benitos (Mexico) and Pinta Island and Marchena Island (Galapagos Islands, Ecuador) (Campbell and Donlan 2005).
Goats were domesticated 10,000 years ago in the highlands of western Iran (Zeder and Hesse 2000, in Campbell and Donlan 2005). Goats are used for their fur and meat as well as for milk and cheese production. Goat meat is the most highly consumed meat source in the world. More goat’s milk is consumed than cow’s milk. Angora goats have long soft fur which is utilised to produce a soft silk-like fabric called mohair. In New Zealand, mohair prices are strong (2006) and farm gate returns are good. Although little is known of optimum farm management systems, it is considered that the farming of angora goats could contribute to the positive growth of the economy (Mohair NZ Business Plan 2006). The French are well known for utilising goat products for economic purposes; making cheese and other goat milk products (Canus Undated). Some cosmetic products have claimed to help eczema sufferers (Johnson 2006).
Uses List
Materials > Skins/leather/fur
Human food and beverage > Fresh meat
Human food and beverage > Milk and dairy products
Prevention and Control
Due to the variable regulations around (de)registration of pesticides, your national list of registered pesticides or relevant authority should be consulted to determine which products are legally allowed for use in your country when considering chemical control. Pesticides should always be used in a lawful manner, consistent with the product's label.
Management Information
Original text compiled by IUCN SSC Invasive Species Specialist Group (ISSG)
Goats have been eradicated from approximately 120 islands and there is hope that native communities will recover rapidly (Hamann 1979, 1993, in Campbell and Donlan 2005). The largest islands on which substantial goat populations have been removed are: Lanai Island (Hawaii), San Clemente (USA), Pinta Island and Santiago Island (Galapagos Islands, Ecuador) and Raoul Island (New Zealand) (Campbell and Donlan 2005; Cruz et al., 2009). In general, goat eradication management for islands larger than 500 hectares requires detailed planning and the use of specialised technology, equipment and personnel. Securing funds for eradication programmes may be an obstacle to goat control (Campbell and Donlan 2005). The recent eradication of feral goats from Santiago Island (58,465 ha) in <5 years cost US$6.1 million, and was achieved with a combination of techniques and technologies (Cruz et al. 2009).
The introduction of modern eradication technology has greatly improved the effectiveness of goat control programmes, making goat eradication more likely. Some control methods include the employment of aerial hunting (helicopter), specially trained goat-hunting dogs, Global Positioning Systems and Geographic Information System techniques as well as sterilised goats marked with radiotelemetry collars, called “Judas goats”, which gravitate to, and therefore detect, wild goat herds. Judas goats are used to find wild herds and are especially suitable for finding the last few survivors or to detect the presence of wild goats when it is uncertain whether they have been eradicated. Hunting dogs are particularly useful in situations when goat density is low and vegetation density is high. Aerial hunting is appropriate in situations where there is less ground cover and a higher density of goats. Eradication is always the better option when compared to short-term control. If short-term control is chosen, goats should be kept at low densities.
Although effective eradication technology is available, Benjamin et al. (2011) demonstrate the potential of strategic water management as a non-lethal technique to protect biodiversity from the impacts of feral goats. In the study area in New South Wales, feral goats were found to remain in close proximity to artificial water points. However, they were able to access and impact upon areas of conservation importance which were nearby to these water points. However, by using fencing, the distance the goats needed to travel from the water points to gain access to conservation areas was increased – this protected biodiversity without eradicating the goat population (although the population was locally reduced). This solution was favourable with local landholders some of whom harvest the feral goats.
Guidelines for managing the impact of feral goats have been developed under the Vertebrate Pest Program (VPP) Australia administered by the Bureau of Resource Sciences (BRS). The purpose of these guidelines is to assist in the development of cost-effective strategies to reduce the damage feral goats cause to production and conservation. Management techniques and strategies for feral goat management are recommended and illustrated by case studies. Deficiencies in knowledge, management and legislation are identified. Please follow this link to view and download Parkes, J., R Henzell & G Pickles, 1995. Managing Vertebrate Pests: Feral Goats
Reproduction
Oestrous cycle
Inter-oestrous interval is 20 days; oestrus duration is 40 hours. Ovulation occurs 30 hours after oestrus begins. Oestrus is seasonal only.
Links to Websites
Name | URL | Comment |
---|---|---|
GISD/IASPMR: Invasive Alien Species Pathway Management Resource and DAISIE European Invasive Alien Species Gateway | https://doi.org/10.5061/dryad.m93f6 | Data source for updated system data added to species habitat list. |
Global register of Introduced and Invasive species (GRIIS) | http://griis.org/ | Data source for updated system data added to species habitat list. |
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US Fish and Wildlife Service, 2010 . 5-Year Review, Short Form Summary: Species Reviewed: Schiedea kealiae (ma oli oli). In: 5-Year Review, Short Form Summary: Species Reviewed: Schiedea kealiae (ma oli oli).US Fish and Wildlife Service . 6 pp.
US Fish and Wildlife Service, 2010 . 5-Year Review, Short Form Summary: Species Reviewed: Schiedea membranacea (no common name). In: 5-Year Review, Short Form Summary: Species Reviewed: Schiedea membranacea (no common name).US Fish and Wildlife Service . 9 pp.
US Fish and Wildlife Service, 2010 . 5-Year Review, Short Form Summary: Species Reviewed: Schiedea stellarioides (no common name). In: 5-Year Review, Short Form Summary: Species Reviewed: Schiedea stellarioides (no common name).US Fish and Wildlife Service . 7 pp.
US Fish and Wildlife Service, 2010 . 5-Year Review, Short Form Summary: Wilkesia hobdyi (dwarf iliau). In: 5-Year Review, Short Form Summary: Wilkesia hobdyi (dwarf iliau).US Fish and Wildlife Service . 8 pp.
US Fish and Wildlife Service, 2010 . 5-year review, Short Form Summary: Zanthoxylum hawaiiense (a'e, manele). In: 5-year review, Short Form Summary: Zanthoxylum hawaiiense (a'e, manele).US Fish and Wildlife Service . 10 pp.
US Fish and Wildlife Service, 2010 . Determination of Endangered Status for 48 Species on Kauai and designation of Critical Habitat: Final Rule. In: Determination of Endangered Status for 48 Species on Kauai and designation of Critical Habitat: Final Rule.US Fish and Wildlife Service . i + 205 pp.
US Fish and Wildlife Service, 2010 . Higo Chumbo (Harrisia portoricensis). 5-Year Review: Summary and Evaluation. In: Higo Chumbo (Harrisia portoricensis). 5-Year Review: Summary and Evaluation . US Fish and Wildlife Service . 19 pp. http://ecos.fws.gov/docs/five_year_review/doc3078.pdf
US Fish and Wildlife Service, 2010 . Plantago princeps (laukahi kuahiwi). 5-Year Review: Summary and Evaluation. In: Plantago princeps (laukahi kuahiwi). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 19 pp.
US Fish and Wildlife Service, 2010 . Recovery Outline for the Kauai Ecosystem. In: Recovery Outline for the Kauai Ecosystem.US Fish and Wildlife Service . 38 pp. + 3 maps.
US Fish and Wildlife Service, 2010 . Schiedea apokremnos (maolioli). 5-Year Review: Summary and Evaluation. In: Schiedea apokremnos (maolioli). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 16 pp.
US Fish and Wildlife Service, 2010 . Sesbania tomentosa (ohai). 5-Year Review: Summary and Evaluation. In: Sesbania tomentosa (ohai). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 24 pp.
US Fish and Wildlife Service, 2010 . Silene lanceolata (no common name). 5-Year Review: Summary and Evaluation. In: Silene lanceolata (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 15 pp.
US Fish and Wildlife Service, 2010 . Spermolepis hawaiiensis (no common name). 5-Year Review: Summary and Evaluation. In: Spermolepis hawaiiensis (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 19 pp.
US Fish and Wildlife Service, 2010 . Tetramolopium filiforme (no common name). 5-Year Review: Summary and Evaluation. In: Tetramolopium filiforme (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 11 pp.
US Fish and Wildlife Service, 2010 . Verbesina dissita (Big-leaved crownbeard). 5-Year Review: Summary and Evaluation. In: Verbesina dissita (Big-leaved crownbeard). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 38 pp. https://ecos.fws.gov/docs/five_year_review/doc3559.pdf
US Fish and Wildlife Service, 2011 . 5-Year Review, Short Form Summary: Species Reviewed: Pritchardia aylmer-robinsonii (wahane). In: 5-Year Review, Short Form Summary: Species Reviewed: Pritchardia aylmer-robinsonii (wahane).US Fish and Wildlife Service . 9 pp.
US Fish and Wildlife Service, 2011 . 5-Year Review, Short Form Summary: Species Reviewed: Pritchardia munroi (lo'ulu). In: 5-Year Review, Short Form Summary: Species Reviewed: Pritchardia munroi (lo'ulu).US Fish and Wildlife Service . 11 pp.
US Fish and Wildlife Service, 2011 . 5-Year Review, Short Form Summary: Vigna o-wahuensis (no common name). In: 5-Year Review, Short Form Summary: Vigna o-wahuensis (no common name).US Fish and Wildlife Service . 10 pp.
US Fish and Wildlife Service, 2011 . Endangered and Threatened Wildlife and Plants; Listing 23 Species on Oahu as Endangered and Designating Critical Habitat for 124 Species. In: Federal Register, 76 ( 148 ) US Fish and Wildlife Service . 46362 - 46593 . http://www.gpo.gov/fdsys/pkg/FR-2011-08-02/pdf/2011-17162.pdf
US Fish and Wildlife Service, 2011 . Hawaiian Dark-rumped Petrel (Pterodroma phaeopygia sandwichensis). 5-Year Review: Summary and Evaluation. In: Hawaiian Dark-rumped Petrel (Pterodroma phaeopygia sandwichensis). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 16 pp.
US Fish and Wildlife Service, 2011 . Isodendrion longifolium (aupaka). 5-Year Review: Summary and Evaluation. In: Isodendrion longifolium (aupaka). 5-Year Review: Summary and Evaluation . US Fish and Wildlife Service . 28 pp. http://ecos.fws.gov/docs/five_year_review/doc3809.pdf
US Fish and Wildlife Service, 2011 . Panicum fauriei var. carteri (no common name). 5-Year Review: Summary and Evaluation. In: Panicum fauriei var. carteri (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 17 pp.
US Fish and Wildlife Service, 2011 . Phyllostegia mannii (no common name). 5-Year Review: Summary and Evaluation. In: Phyllostegia mannii (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 14 pp.
US Fish and Wildlife Service, 2011 . Schiedea haleakalensis (no common name). 5-Year Review: Summary and Evaluation. In: Schiedea haleakalensis (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 17 pp.
US Fish and Wildlife Service, 2011 . Schiedea hookeri (no common name). 5-Year Review: Summary and Evaluation. In: Schiedea hookeri (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 20 pp.
US Fish and Wildlife Service, 2011 . Schiedea lydgatei (no common name). 5-Year Review: Summary and Evaluation. In: Schiedea lydgatei (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 17 pp.
US Fish and Wildlife Service, 2011 . Schiedea sarmentosa (no common name). 5-Year Review: Summary and Evaluation. In: Schiedea sarmentosa (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 14 pp.
US Fish and Wildlife Service, 2011 . Urera kaalae (opuhe). 5-Year Review: Summary and Evaluation. In: Urera kaalae (opuhe). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 19 pp.
US Fish and Wildlife Service, 2012 . Endangered and Threatened Wildlife and Plants; Endangered Status for 23 Species on Oahu and Designation of Critical Habitat for 124 Species; Final Rule. In: Federal Register, 77 ( 181 ) US Fish and Wildlife Service . 57648 - 57862 . https://www.gpo.gov/fdsys/pkg/FR-2012-09-18/pdf/2012-19561.pdf
US Fish and Wildlife Service, 2012 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Synthliboramphus hypoleucus. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Synthliboramphus hypoleucus . US Fish and Wildlife Service . 24 pp. http://ecos.fws.gov/docs/candidate/assessments/2012/r8/B098_V01.pdf
US Fish and Wildlife Service, 2013 . Endangered and Threatened Wildlife and Plants; Determination of Endangered Species Status for 15 Species on Hawaii Island; Final Rule. In: Federal Register, 78 ( 209 ) US Fish and Wildlife Service . 64638 - 64690 . https://www.gpo.gov/fdsys/pkg/FR-2013-10-29/pdf/2013-24103.pdf
US Fish and Wildlife Service, 2013 . Endangered and Threatened Wildlife and Plants; Determination of Endangered Status for 38 Species on Molokai, Lanai, and Maui; Final Rule. In: Federal Register, 78 ( 102 ) US Fish and Wildlife Service . 32014 - 32065 . https://www.gpo.gov/fdsys/pkg/FR-2013-05-28/pdf/2013-12105.pdf
US Fish and Wildlife Service, 2013 . Endangered and Threatened Wildlife and Plants; Review of Native Species That are Candidates for Listing as Endangered or Threatened; Annual Notice of Findings on Resubmitted Petitions; Annual Description of Progress on Listing Actions. In: Federal Register, 78 ( 226 ) US Fish and Wildlife Service . 70104 - 70162 . https://www.gpo.gov/fdsys/pkg/FR-2013-11-22/pdf/2013-27391.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Gardenia remyi. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Gardenia remyi . US Fish and Wildlife Service . 17 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/Q33N_P01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Huperzia (= Phlegmariurus) stemmermanniae. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Huperzia (= Phlegmariurus) stemmermanniae . US Fish and Wildlife Service . 14 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/S02B_P01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus assimulans. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus assimulans . US Fish and Wildlife Service . 33 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/I0GQ_I01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus facilis. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus facilis . US Fish and Wildlife Service . 32 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/I0GY_I01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus hilaris. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus hilaris . US Fish and Wildlife Service . 31 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/I0HT_I01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus kuakea. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus kuakea . US Fish and Wildlife Service . 29 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/I0VM_I01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus longiceps. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus longiceps . US Fish and Wildlife Service . 33 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/I0HC_I01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus mana. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus mana . US Fish and Wildlife Service . 30 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/I0VL_I01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Joinvillea ascendens ascendens. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Joinvillea ascendens ascendens . US Fish and Wildlife Service . 22 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/Q1XN_P01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Ochrosia haleakalae. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Ochrosia haleakalae . US Fish and Wildlife Service . 17 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/Q3ES_P01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Pseudognaphalium sandwicensium var. molokaiense. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Pseudognaphalium sandwicensium var. molokaiense . US Fish and Wildlife Service . 13 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/Q1WN_P01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Ranunculus hawaiensis. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Ranunculus hawaiensis . US Fish and Wildlife Service . 16 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/Q38H_P01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Schiedea pubescens. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Schiedea pubescens . US Fish and Wildlife Service . 14 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/Q2VZ_P01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Streptanthus bracteatus. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Streptanthus bracteatus . US Fish and Wildlife Service . 24 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r2/Q1R7_P01.pdf
US Fish and Wildlife Service, 2015 . Endangered and Threatened Wildlife and Plants; Endangered Status for 16 Species and Threatened Status for 7 Species in Micronesia. In: Federal Register, 80 ( 190 ) US Fish and Wildlife Service . 59424 - 59497 . https://www.gpo.gov/fdsys/pkg/FR-2015-10-01/pdf/2015-24443.pdf
US Fish and Wildlife Service, 2015 . Endangered and Threatened Wildlife and Plants; Endangered Status for 49 Species From the Hawaiian Islands. In: Federal Register, 80 ( 189 ) US Fish and Wildlife Service . 58820 - 58909 . https://www.gpo.gov/fdsys/pkg/FR-2015-09-30/pdf/2015-24305.pdf
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