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15 September 2010

Capra hircus (goats)

Datasheet Types: Natural enemy, Livestock species, Invasive species, Host animal

Abstract

This datasheet on Capra hircus covers Identity, Overview, Distribution, Dispersal, Biology & Ecology, Physiology, Impacts, Uses, Prevention/Control, Management, Further Information.

Identity

Preferred Scientific Name
Capra hircus Linnaeus 1758
Preferred Common Name
goats
Other Scientific Names
Capra aegagrus hircus
Capra indicus
International Common Names
English
billy
goat
kid
kids
nanny
Spanish
cabras
French
chèvres
Local Common Names
Germany
Hausziege
Ziegen
Italy
capre
Portugal
cabras

Pictures

Capra hircus (goats); goats grazing in an Argan tree (Argania spinosa). Essaouira, Morocco. June 2013.
Goats grazing in a tree
Capra hircus (goats); goats grazing in an Argan tree (Argania spinosa). Essaouira, Morocco. June 2013.
©Grand Parc - Bordeaux, France/via wikipedia - CC BY 2.0
Capra hircus (goats); herd of Saanen dairy goats. Lukoz Farm, Russia. September 2008.
Herd of dairy goats
Capra hircus (goats); herd of Saanen dairy goats. Lukoz Farm, Russia. September 2008.
©Efras/via wikipedia - CC BY-SA 3.0
Capra hircus (goats); herd of Tibetan goats, with their goatherd. Tibet. October 2005.
Herd of Tibetan goats
Capra hircus (goats); herd of Tibetan goats, with their goatherd. Tibet. October 2005.
©Adam Schorsch/via wikipedia - CC BY 2.0
Capra hircus (goats); female Toggenburg goat from "Zlatusha" farm of the Institute of Animal Science, Kostinbrod, Bulgaria, 1995.
Female
Capra hircus (goats); female Toggenburg goat from "Zlatusha" farm of the Institute of Animal Science, Kostinbrod, Bulgaria, 1995.
©Nadezhda Petrova

Summary of Invasiveness

The goat ( Capra hircus ) was domesticated 10,000 years ago in the highlands of western Iran. These herbivores have a highly varied diet and are able to utilise a larger number of plant species than other livestock. Goats alter plant communities and forest structure and threaten vulnerable plant species. The reduction of vegetation reduces shelter options for native animals and overgrazing in native communities leads to ecosystem degradation. Feral goats spread disease to native animals. Native fauna on islands are particularly susceptible.

Taxonomic Tree

This content is currently unavailable.

Breeds

Breed
Abaza goats
Abgal goats
Abyssinian Short-eared goats
Adal goats
Adamello Blond goats
Adany goats
African Pygmy goats
Algarvia goats
Alpine goats (Europe, S America)
Altai Mountain goats
Anatolian Black goats
Anglo-Nubian goats
Angora goats
Appenzell goats
Aquila goats
Arabia goats
Arapawa goats
Askanian mohair goats
Asmari goats
Aswad goats
Attaouia goats
Auckland Island goats
Australian feral goats
Azerbaijan goats
Bach Thao goats
Bae goats
Bagot goats
Baigani goats
Baladi goats
Baltistani goats
Banjiao goats
Bantu goats
Barbari goat
Barki goats
Beetal goat (Beetal)
Beiari goats
Belgian Fawn goats
Belgian Saanen goats
Benadir goats
Benevento goats
Bengal goat
Berber goats
Bermeya goats
Bhuj goats
Bimal goats
Boer goats
Bornu White goats
British Alpine goats
Buchi goats
Bugi Tori goats
Bugri goats
Bukovica goats
Burmese goats
Camoschiata delle Alpi (goat)
Campobasso goats
Caninde goats
Capore goats
Capra aegagrus
Capra ibex sibirica
Carpathian goats
Catalan goats
Central Asian Local Coarse-haird goats
Cevennes White goats
Chamois Coloured goats
Changthangi goats
Chappar goats
Charnequeira goats
Chengde Polled goats
Chengdu Brown goats
Chigu goats
Cilento Black goats
Cilento Fawn goats
Cilento Grey goats
Co goats
Congo Dwarf goats
Corsican goats
Cosenza goats
Cou-Clair de Berry goats
Creole goats
Crimean goats
Criollo goats
Crioulo goats
Cyprus goats
Czech Brown Short-haired goats
Czech White Short-haired goats
Dagestan goats
Dagestan White goats
Dairy goats
Dalua goats
Damagaran Dapple-grey goats
Damani goats
Damara goats
Damascus goats
Danish Landrace goats
Dera Din Panah goats
Desi goats
Don goats
Don-Kirgiz cross
Dragobia goats
Duan goats
Dutch Dwarf goats
Dutch Landrace goats
Dutch Pied goats
Erzgebirge goats
Estonian goats
Etna Silver goats
Fiji goats
Finnish Landrace goats
Florida Sevillana goats
French Alpine goats
Fuqing goats
Gaddi goats
Galapagos goats
Gallega goats
Ganjam goats
Garganica goats
Garre goats
German Improved Fawn goats
Germasia goats
Girgentana goats
Gobi Wool goats
Gohilwadi goats
Goingeget goats
Golden Guernsey goats
Gorki goats
Granada goat
Greek indigenous goats
Grey Molise goats
Grisons Striped goats
Guadalupe Island goats
Guadarrama goats
Guanzhong Dairy goats
Guizhou White goats
Gwembe goats
Hailun goats
Haimen goats
Harz goats
Hasi goats
Hebei dairy goats
Heilongjian Dairy goats
Hejazi goats
Hexi Cashmere goats
Huanghuai goats
Icelandic goats
Indian Mohair goats
Indo-Chinese goats
Ingessana goats
Ionica goats
Iraqi goats
Irish goats
Israeli Saanen goats
Istrian goats
Jabaly goats
Jamnapari goat
Jarakheil goats
Jattal goats
Jhakrana goats
Jining Grey goats
Juan Fernandez goats
Jurdana goats
Kabyle goats
Kaghani goats
Kajli goats
Kamori goats
Kannaiadu
Kano Brown goats
Karachai goats
Katjang goats
Katsina Light-brown goats
Kazakh goats
Khurasani goats
Kigezi goats
Kiko goats
Kilis goats
Kinder goats
Kirgiz goats
Kohai Ghizer goats
Kooti goats
Kottukachchiya goats
Kurdi goats
Kutchi goats
Labi goats
Labri goats
Lamancha goats
Laoshan Dairy goats
Lehri goats
Leizhou goats
Liaoning Cashmere goats
Libyan goats
Liqenasi goats
Lithuanian Native goats
Longlin goats
Lori goats
Macedonian goats
Machaeras goats
Malabari goats
Malawian goats
Maltese goats
Mambilla goats
Maradi goats
markhor goats
Marota goats
Marungu goats
Marwari goat
Massif Central goats
Mati goats
Matou goats
Maure goats
Mawr goats
Mehsana goats
Meridional goats
Merturi goats
Mingrelian goats
Miriz goats
Moldavian goats
Mongolian goats
Montecristo goats
Moroccan Black goats
Moussoro goats
Moxotó goats
Mubende goats
Murcia-Granada goat
Murme e Zeze goats
Muzhake goats
Myotonic goats
Mzabite goats
Nachi goats
Nadjdi goats
Ndebele goats
Neapolitan goats
Negev goats
Nepal Hill goats
New Zealand Base Stock goats
Nigerian Dwarf goats
Nigerian goats
Nilotic goats
North Russian goats
Norwegian goats
Nubisk goats
Oberhasli goats
Ogaden goats
Okinawa goats
Old English goats
Orenburg goats
Orobica goats
Osmanabadi goats
Ovambo goats
Pafuri goats
Pateri goats
Peacock goats
Peratiki goats
Philippine goats
Piamiri goats
Pinzgau goats
Poitou goat
Polish Improved Fawn goats
Pomellata goats
Portuguese ibex
Potentina goats
Potenza goats
Pothohari goats
Provencal goats
Pygora goats
Pyrenean goats
Rahnama goats
Raini goats
Ramdhan goats
Red Bosnian goats
Red Mediterranean goats
Red Sokoto goats
Repartida goats
Roccaverano goats
Rock Alpine goats
Rove goats
Rovmit goats
Saanen goats
Sahelian goat
Saidi goat
Samar goats
San Clemente goats
Sangamneri goats
Santa Catalina goats
Sardinian goats
Screziata goats
Sempione goats
Serpentina goats
Serrana goats
Shannan White goats
Sharkawi goats
Shiba goats
Shukria goats
Shurri goats
Sinai goats
Sinazongwe goats
Sind wild goat
Sindhal goats
Sirli goats
Sirohi goats
Skopelos goats
Slovakian Brown Short-haired goats
Small East African goat
Somali Arab goats
Somali goats
Southern Sudan goats
Soviet Mohair goats
Spanish goats
SRD goats
Sri Lankan goats
Stiefelgeiss (goat)
Sudanese Desert goats
Sudanese Nubian goats
Sundgau goats
Surdudi goats
Surti goats
Swedish Landrace goats
Syrian Mountain goats
Syrian type goats
Taihang goats
Taiwan goats
Taiz Black goats
Taiz Red goats
Tajik goats
Tali goats
Tarai goats
Tauern Pied goats
Teddy goats
Teramo goats
Thai goats
Thuringian goats
Tibetan goats
Tihami goats
Toggenburg goat
Tokara goats
Toposa goats
Tranga goats
Tremiti goats
Tswana goats
Tuareg goats
Tuni goats
Turkmen goats
Unjuul goats
Utegangsgeit goats
Uuliin Bor goats
Uzbek Black goats
Uzbek goats
Val di Livo goats
Valais Blackneck goats
Valfortorina goats
Vatani goats
Velipoja goats
Vendi goats
Verzasca goats
Volgograd White goats
Voltaïque goats
Wahati goats
Wallische geit (goat)
Welsh goats
West African Dwarf goats
White Swiss Improved goats
Wuan goats
Xinjiang goats
Xuhai goats
Yanbian Dairy goats
Yei goats
Yemen Mountain goats
Yemso goats
Yugoslav Mountainous goats
Yugoslav Saanen goats
Zalawadi goats
Zaraibi goat
Zhiwulin Black goats
Zhongwei goats
Zwergziege (goat)

Description

Males weigh between 45 and 55 kilos and females weigh between 25 and 35 kilos. Colouration is highly variable from mostly black, to various shades of brown, and from single-coloured to multi-coloured. Black anterior with brown posterior is a common pattern. Horns are dimorphic, having homonymous spiral and anterior keel. Males are bearded and produce pheromones during the breeding season.

Pathogens Carried

This content is currently unavailable.

Distribution

Goats were domesticated 10,000 years ago in the highlands of western Iran. Feral goats are derived from domestic goats, originally native to Asia.

Distribution Map

This content is currently unavailable.

Distribution Table

This content is currently unavailable.

Means of Movement and Dispersal

Introduction pathways to new locations
Other: Goats ( Capra hircus ) were released as potential food for people marooned by shipwrecks.
Transportation of domesticated animals: Goats ( Capra hircus ) were introduced to islands for their milk.

Local dispersal methods
Escape from confinement:
Natural dispersal (local):

Pathway Causes

Habitat

Goats usually move in herds that roam over territories up to 20km wide. Sometimes herd ranges can be as small as 100m 2 . Males usually wander more widely than females. Grasslands, scrub lands, rocky outcrops and semi-open or open forests are all used extensively by goats as habitat substrate. In bad weather, they may seek shelter under rocky ledges (SPREP, 2000).

Habitat List

CategorySub categoryHabitatPresenceStatus
TerrestrialTerrestrial – ManagedManaged forests, plantations and orchardsPresent, no further detailsHarmful (pest or invasive)
TerrestrialTerrestrial ‑ Natural / Semi-naturalNatural forestsPresent, no further detailsHarmful (pest or invasive)
TerrestrialTerrestrial ‑ Natural / Semi-naturalNatural grasslandsPresent, no further detailsHarmful (pest or invasive)
TerrestrialTerrestrial ‑ Natural / Semi-naturalScrub / shrublandsPresent, no further detailsHarmful (pest or invasive)

Biology and Ecology

Nutrition
Goats are herbivores and will forage on any palatable plants in their home range. Goats have rather large rumino-reticular volume so they are able to subsist on poorer quality plants than most herbivores, therefore goats can survive and subsist in heavily exploited environments.
In a study conducted by Chimera, Coleman and Parkes (1995) the rumen contents of 49 goats (captured in 1989) were identified and the dry weight of each component was measured to produce a breakdown of the diet of a small, unique remnant population (now extinct) of feral goats on Auckland Island (a Sub-Antarctic island located south of New Zealand). Woody plants and grasses made up the bulk of their diet (41% and 39%, respectively); seaweeds made up 13%, ferns 4% and herbaceous species 3%. At least 40 plant species were eaten by the goats, but just three species made up half of the total: rata ( Metrosideros umbellate ), snow tussock ( Chionochloa Antarctica ) and kelp ( Durvillea Antarctica ). Rata ( Metrosideros umbellate ), Coprosma foetidissima, Pseudopanax simplex and Carex appressa were the most commonly found species - eaten in at least some quantity by 90%, 80%, 76% and 69% of the goats respectively.
Unidentified grasses composed 21.4% of the total contents (dry weight) and were found in 100% of goats. NB: This population had a unique genetic makeup as it evolved separately for over 100 years on an isolated island with a cold and harsh climate and so it may not represent the dietary preferences of all goats. However, it highlights the huge adaptability of goats and their ability to subsist in particular and inhospitable landscapes.
Reproduction
Both sexes are physiologically capable of reproduction at about 6 months of age. Dominant males fight to win females. They follow a serial pattern and attend to one female after another as they come into oestrus. In one sense this is termed polygyny, as males breed with as many females as they can during a breeding period, but more properly this is serial monogamy as a male will tend a female for extended periods, both before and after copulation and before leaving in search of a different mate.
Realised reproduction varies among populations, seasons and years. At best a typical female goat would produce one young in its first pregnancy and twin kids in subsequent pregnancies. However, most females are at less than optimal condition, which may result in a twinning rate of as little as 0%. At best, herds may produce twins at a rate of 80% or more. Triplets are not uncommon.

Physiology Table

TypeValue
Heart rate (beats per minute)70 - 135
Daily urine volume (mL/kg of body weight)10 - 40
Rectal temp (?C+0.5?C)39

Haematology

TypeValue
Haemoglobin (g/dL)8 - 12
PCV (%)22 - 38
RBC (x10 6 /µL)8 - 18
MCV (fL)16 - 25
MCH (pg)5 - 8
MCHC (g/dL)30 - 36
Reticulocytes (% of RBC)0
WBC (/µL)4000 - 13000
Neutrophils (mature) (/µL)1200 - 7200
Lymphocytes (/µL)2000 - 9000
Monocytes (/µL)0 - 550
Eosinophils (/µL)50 - 650
Basophils (/µL)0 - 120
Platelets (x10 5 /µL)3 - 6
Plasma proteins (g/dL)6 - 7.5
Fibrinogen (mg/dL)100 - 400

Blood Chemistry

TypeValue
Alkaline phosphatase (U/L)42 - 775
Aspartate aminotranferase (U/L)67 - 117
Bicarbonate (mmol/L)26 - 30
Bilirubin - total (mg/dL)0 - 0.1
Calcium (mg/dL)8.5 - 10.2
Chloride (mEq/L)99 - 110
Cholesterol (mg/dL)80 - 130
γ-glutamyltransferase (U/L)43 - 71
Glucose (mg/dL)58 - 76
Magnesium (mg/dL)2.8 - 3.6
Potassium (mEq/L)3.5 - 6.7
Sodium (mEq/L)142 - 155
Sorbitol dehydrogenase (U/L)35 - 80
Urea Nitrogen (mg/dL)15 - 33
Total Protein (g/dL)5.9 - 7.4
Albumin (g/dL)2.7 - 3.9
a1 globulin (g/dL)0.5 - 0.7
β1 globulin (g/dL)0.7 - 1.2
β2 globulin (g/dL)0.3 - 0.6
γ1 globulin (g/dL)0.9 - 3

List of Diseases and Disorders

This content is currently unavailable.

Natural enemy of

This content is currently unavailable.

Impact Summary

CategoryImpact
Biodiversity (generally)Negative
Environment (generally)Negative
Livestock productionNegative

Impact

General Impacts

Original text compiled by IUCN SSC Invasive Species Specialist Group (ISSG)

  Biodiversity on islands is vulnerable to introduced species, and the vegetation tends to lack adaptions to mammalian herbivores (Campbell and Donlan 2005). Unfortunately, goats (

Capra hircus

) have been established on many islands. Goats alter plant communities and forest structures and threaten vulnerable plant species; the flow-down effect of these outcomes includes increased soil erosion and the reduction of native fauna that share a similar environmental niche (Spatz and Mueller-Dombois 1973, Coblentz 1978, Parkes 1984, Brennan 1986, Coblentz and Van Vuren 1987, Cronk 1989, Walker 1991, Moran 1996, Desender

et al

. 1999, in Campbell and Donlan 2005). In some island ecosystems it has been the case that goats are the most destructive herbivore present (King, 1985). Feral goats are particularly destructive in such environments and cause a huge loss of native vegetation due to their grazing habits. This leads to ecosystem degradation and biodiversity loss (Coblentz 1978; Schofield 1989; Moran 1996; Desender

et al.

1999 in Campbell and Donlan 2005).

Threatened Species

Threatened speciesWhere threatenedMechanismsReferencesNotes
Acrocephalus luscinia (nightingale reed warbler)
Guam
Northern Mariana Islands
Herbivory/grazing/browsing
 
Adelocosa anops (Kauai cave wolf spider)
Hawaii
Herbivory/grazing/browsing
 
Bidens amplectens (koko'olau)
Hawaii
Herbivory/grazing/browsing
 
Bidens campylotheca subsp. waihoiensis (ko`oko`olau)
Hawaii
Herbivory/grazing/browsing
 
Bidens conjuncta (ko`oko`olau)
Hawaii
Herbivory/grazing/browsing
 
Bidens micrantha subsp. ctenophylla (ko`oko`olau)
Hawaii
Herbivory/grazing/browsing
 
Calamagrostis hillebrandii (Hillegrands reedgrass)
Hawaii
Herbivory/grazing/browsing
 
Canavalia pubescens (jack bean)
Hawaii
Herbivory/grazing/browsing
 
Cinclodes aricomae
Peru
Herbivory/grazing/browsing
 
Cyanea asplenifolia (haha)
Hawaii
Herbivory/grazing/browsing
 
Cyanea calycina
Hawaii
Herbivory/grazing/browsing
 
Cyanea duvalliorum
Hawaii
Herbivory/grazing/browsing
 
Cyanea horrida (prickly cyanea)
Hawaii
Herbivory/grazing/browsing
 
Cyanea kunthiana (Kunth's cyanea)
Hawaii
Herbivory/grazing/browsing
 
Cyanea lanceolata (lanceleaf cyanea)
Hawaii
Herbivory/grazing/browsing
 
Cyanea magnicalyx
Hawaii
Herbivory/grazing/browsing
 
Cyanea maritae
Hawaii
Herbivory/grazing/browsing
 
Cyanea marksii (Marks' cyanea)
Hawaii
Herbivory/grazing/browsing
 
Cyanea mauiensis
Hawaii
Herbivory/grazing/browsing
 
Cyanea munroi
Hawaii
Herbivory/grazing/browsing
 
Cyanea obtusa (bluntlobe cyanea)
Hawaii
Herbivory/grazing/browsing
 
Cyanea profuga (Mapulehu Valley cyanea)
Hawaii
Herbivory/grazing/browsing
 
Cyanea recta (Kealia cyanea)
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Cyanea solanacea (popolo)
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Cyclura stejnegeri (Mona Island iguana)
Puerto Rico
Competition - monopolizing resources
 
Cyrtandra ferripilosa
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Cyrtandra filipes
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Cyrtandra limahuliensis (Limahuli cyrtandra)
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Cyrtandra oxybapha
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Doryopteris takeuchii (Takeuch's lipfern)
Hawaii
Herbivory/grazing/browsing
 
Dracaena forbesii (hala pepe)
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Chrysodracon hawaiiensis (hala pepe)
Hawaii
Herbivory/grazing/browsing
 
Drosophila aglaia
Hawaii
Herbivory/grazing/browsing
 
Drosophila differens (Hawaiian picture-wing fly)
Hawaii
Herbivory/grazing/browsing
 
Drosophila digressa
Hawaii
Herbivory/grazing/browsing
 
Drosophila hemipeza
Hawaii
Herbivory/grazing/browsing
 
Drosophila heteroneura
Hawaii
Herbivory/grazing/browsing
 
Drosophila montgomeryi
Hawaii
Herbivory/grazing/browsing
 
Drosophila musaphilia
Hawaii
Herbivory/grazing/browsing
 
Drosophila ochrobasis
Hawaii
Herbivory/grazing/browsing
 
Drosophila substenoptera
Hawaii
Herbivory/grazing/browsing
 
Emballonura semicaudata rotensis (Pacific sheath-tailed bat)
Northern Mariana Islands
Ecosystem change / habitat alteration
 
Epicrates monensis monensis (Mona boa)
Puerto Rico
Ecosystem change / habitat alteration
 
Festuca hawaiiensis (Hawai'i fescue)
Hawaii
Herbivory/grazing/browsing
Ecosystem change / habitat alteration
 
Festuca molokaiensis
Hawaii
Herbivory/grazing/browsing
Ecosystem change / habitat alteration
 
Gardenia remyi (Remy's gardenia)
Hawaii
Herbivory/grazing/browsing
Ecosystem change / habitat alteration
 
Harrisia portoricensis (higo chumbo)
Puerto Rico
Herbivory/grazing/browsing
 
Huperzia stemmermanniae (Stemmermann's clubmoss)
Hawaii
Herbivory/grazing/browsing
Ecosystem change / habitat alteration
 
Hylaeus assimulans (assimulans yellow-faced bee)
Hawaii
Herbivory/grazing/browsing
Ecosystem change / habitat alteration
 
Hylaeus facilis (easy yellow-faced bee)
Hawaii
Herbivory/grazing/browsing
Ecosystem change / habitat alteration
 
Hylaeus hilaris (hilaris yellow-faced bee)
Hawaii
Herbivory/grazing/browsing
Ecosystem change / habitat alteration
 
Hylaeus kuakea (Hawaiian yellow-faced bee)
Hawaii
Herbivory/grazing/browsing
Ecosystem change / habitat alteration
 
Hylaeus longiceps
Hawaii
Herbivory/grazing/browsing
Ecosystem change / habitat alteration
 
Hylaeus mana
Hawaii
Herbivory/grazing/browsing
Ecosystem change / habitat alteration
 
Isodendrion longifolium (longleaf isodendrion)
Hawaii
Herbivory/grazing/browsing
Ecosystem change / habitat alteration
 
Joinvillea ascendens ascendens
Hawaii
Herbivory/grazing/browsing
Ecosystem change / habitat alteration
 
Korthalsella taenioides (Degener's korthal mistletoe)
Hawaii
Herbivory/grazing/browsing
Ecosystem change / habitat alteration
 
Leiolopisma telfairii (Round Island skink)
Mauritius
Ecosystem change / habitat alteration
 
Lobelia monostachya (Waianae Range lobelia)
Hawaii
Herbivory/grazing/browsing
 
Lobelia niihauensis (Niihau lobelia)
Hawaii
Herbivory/grazing/browsing
 
Loxioides bailleui (palila)
Hawaii
Herbivory/grazing/browsing
 
Melanthera tenuifolia (Waianae Range nehe)
Hawaii
Herbivory/grazing/browsing
 
Melanthera venosa (spreading nehe)
Hawaii
Herbivory/grazing/browsing
 
Melanthera waimeaensis (Waimea Canyon nehe)
Hawaii
Herbivory/grazing/browsing
 
Melicope christophersenii
Hawaii
Herbivory/grazing/browsing
Ecosystem change / habitat alteration
 
Melicope hiiakae
Hawaii
Herbivory/grazing/browsing
Ecosystem change / habitat alteration
 
Melicope makahae
Hawaii
Ecosystem change / habitat alteration
 
Melicope makahae
Hawaii
Ecosystem change / habitat alteration
 
Mucuna sloanei var. persericea
Hawaii
Herbivory/grazing/browsing
Ecosystem change / habitat alteration
 
Myrsine linearifolia (narrowleaf colicwood)
Hawaii
Competition - monopolizing resources
 
Nothocestrum latifolium (broadleaf aiea)
Hawaii
Herbivory/grazing/browsing
Ecosystem change / habitat alteration
 
Nothocestrum peltatum (Oahu aiea)
Hawaii
Herbivory/grazing/browsing
 
Nototrichium humile (kaala rockwort)
Hawaii
Herbivory/grazing/browsing
 
Oceanodroma castro (band-rumped storm-petrel)
Montana
Ecosystem change / habitat alteration
 
Ochrosia haleakalae (island yellowwood)
Hawaii
Herbivory/grazing/browsing
Ecosystem change / habitat alteration
 
Ochrosia kilaueaensis (Hawai'i yellowwood)
Hawaii
Herbivory/grazing/browsing
 
Oreomystis bairdi (akikiki)
Hawaii
Herbivory/grazing/browsing
 
Oreomystis mana (Hawaii creeper)
Hawaii
Herbivory/grazing/browsing
 
Osmoxylon mariannense
Northern Mariana Islands
Herbivory/grazing/browsing
 
Ovis ammon (argali)
Kyrgyzstan
Mongolia
Tajikistan
Competition - monopolizing resources
 
Ovis canadensis sierrae (Sierra Nevada bighorn sheep)
California
Pathogenic
 
Panicum fauriei (Carter's panicgrass)
Hawaii
Ecosystem change / habitat alteration
 
Paroreomyza flammea (Molokai creeper)
Hawaii
Ecosystem change / habitat alteration
 
Partula gibba (humped tree snail)
Guam
Northern Mariana Islands
Ecosystem change / habitat alteration
 
Partula langfordi (Langford's tree snail)
Northern Mariana Islands
Ecosystem change / habitat alteration
 
Partula radiolata (Guam tree snail)
Guam
Ecosystem change / habitat alteration
 
Partulina semicarinata (Lanai tree snail)
Hawaii
Ecosystem change / habitat alteration
 
Partulina variabilis (Lanai tree snail)
Hawaii
Ecosystem change / habitat alteration
 
Peucedanum sandwicense (makou)
Hawaii
Herbivory/grazing/browsing
 
Phyllostegia glabra var. lanaiensis (ulihi phyllostegia)
Hawaii
Ecosystem change / habitat alteration
 
Phyllostegia knudsenii (Waimea phyllostegia)
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Phyllostegia mannii (Mann's phyllostegia)
Hawaii
Ecosystem change / habitat alteration
 
Phyllostegia mollis (Waianae Range phyllostegia)
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Phyllostegia parviflora (smallflower phyllostegia)
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Phyllostegia renovans (red-leaf phyllostegia)
Hawaii
Herbivory/grazing/browsing
 
Phyllostegia waimeae (Kauai phyllostegia)
Hawaii
Herbivory/grazing/browsing
 
Phyllostegia wawrana (fuzzystem phyllostegia)
Hawaii
Herbivory/grazing/browsing
 
Phytotoma raimondii (Peruvian plantcutter)
Peru
Ecosystem change / habitat alteration
 
Pittosporum napaliense (royal cheesewood)
Hawaii
Herbivory/grazing/browsing
 
Plantago hawaiensis (Hawai'i plantain)
Hawaii
Herbivory/grazing/browsing
 
Plantago princeps
Hawaii
Herbivory/grazing/browsing
 
Platydesma cornuta var. cornuta
Hawaii
Herbivory/grazing/browsing
Ecosystem change / habitat alteration
 
Platydesma cornuta var. decurrens
Hawaii
Herbivory/grazing/browsing
Ecosystem change / habitat alteration
 
Platydesma rostrata
Hawaii
Herbivory/grazing/browsing
Ecosystem change / habitat alteration
 
Poa mannii (Mann's bluegrass)
Hawaii
Herbivory/grazing/browsing
 
Portulaca sclerocarpa
Hawaii
Herbivory/grazing/browsing
 
Pritchardia hardyi (Makaleha pritchardia)
Hawaii
Herbivory/grazing/browsing
 
Pritchardia kaalae (Waianae Range pritchardia)
Hawaii
Herbivory/grazing/browsing
 
Pritchardia munroi (Kamalo pritchardia)
Hawaii
Herbivory/grazing/browsing
 
Pritchardia napaliensis
Hawaii
Herbivory/grazing/browsing
 
Pritchardia remota (Remota loula palm)
Hawaii
Herbivory/grazing/browsing
 
Pseudognaphalium sandwicensium var. molokaiense (enaena)
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Psittirostra psittacea (Ou)
Hawaii
Herbivory/grazing/browsing
 
Psychotria grandiflora (large-flowered balsamo)
Hawaii
Herbivory/grazing/browsing
 
Psychotria hobdyi (Hobdy's wild-coffee)
Hawaii
Herbivory/grazing/browsing
 
Pteralyxia kauaiensis (Kauai pteralyxia)
Hawaii
Herbivory/grazing/browsing
 
Pteralyxia laurifolia (ridged pteralyxia)
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Pterodroma sandwichensis
Hawaii
Herbivory/grazing/browsing
 
Pteropus mariannus mariannus (Mariana fruit bat)
Guam
Northern Mariana Islands
Ecosystem change / habitat alteration
 
Ranunculus hawaiensis (Hawai'ian buttercup)
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Ranunculus mauiensis (Maui buttercup)
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Remya kauaiensis (Kauai remya)
Hawaii
Ecosystem change / habitat alteration
 
Remya mauiensis (Maui remya)
Hawaii
Competition (unspecified)
 
Remya montgomeryi (Kalalau Valley remya)
Hawaii
Ecosystem change / habitat alteration
 
Sanicula mariversa (Waianae Range blacksnakeroot)
Hawaii
Ecosystem change / habitat alteration
 
Schiedea apokremnos (Kauai schiedea)
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Schiedea haleakalensis (Haleakala schiedea)
Hawaii
Ecosystem change / habitat alteration
 
Schiedea helleri (Kaholuamanu schiedea)
Hawaii
Ecosystem change / habitat alteration
 
Schiedea helleri (Kaholuamanu schiedea)
Hawaii
Ecosystem change / habitat alteration
 
Schiedea hookeri (sprawling schiedea)
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Schiedea jacobii (Jacobi's schiedea)
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Schiedea kaalae (Oahu schiedea)
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Schiedea kauaiensis (Kauai schiedea)
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Schiedea kealiae (Waianae Range schiedea)
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Schiedea lydgatei (Kamalo Gulch schiedea)
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Schiedea membranacea
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Schiedea nuttallii
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Schiedea pubescens
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Schiedea salicaria
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Schiedea sarmentosa
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Schiedea spergulina var. leiopoda
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Schiedea spergulina var. spergulina
Hawaii
Herbivory/grazing/browsing
 
Schiedea stellarioides
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Sesbania tomentosa
Hawaii
Ecosystem change / habitat alteration
 
Sibara filifolia (Santra Cruz Island Rockcress)
California
Herbivory/grazing/browsing
 
Silene alexandri
Hawaii
Ecosystem change / habitat alteration
 
Silene hawaiiensis (Hawaii catchfly)
Hawaii
Herbivory/grazing/browsing
 
Silene lanceolata (Kauai catchfly)
Hawaii
Ecosystem change / habitat alteration
 
Solanum incompletum
Hawaii
Ecosystem change / habitat alteration
 
Solanum sandwicense
Hawaii
Ecosystem change / habitat alteration
 
Spermolepis hawaiiensis (Hawaii scaleseed)
Hawaii
Ecosystem change / habitat alteration
 
Stenogyne campanulata (Kalalau Valley stenogyne)
Hawaii
Herbivory/grazing/browsing
 
Stenogyne kanehoana (Oahu stenogyne)
Hawaii
Herbivory/grazing/browsing
 
Stenogyne purpurea (purplefruit stenogyne)
Hawaii
Herbivory/grazing/browsing
 
Streptanthus bracteatus (bracted twistflower)
Texas
Herbivory/grazing/browsing
 
Synthliboramphus hypoleucus (Xantus' murrelet)
Canada
Mexico
California
Oregon
Ecosystem change / habitat alteration
 
Tetramolopium arenarium (Maui tetramolopium)
Hawaii
Herbivory/grazing/browsing
 
Tetramolopium filiforme (ridgetop tetramolopium)
Hawaii
Herbivory/grazing/browsing
 
Tetramolopium lepidotum (Waianae Range tetramolopium)
Hawaii
Herbivory/grazing/browsing
 
Tetramolopium remyi (Awalua Ridge tetramolopium)
Hawaii
Herbivory/grazing/browsing
 
Tetraplasandra flynnii
Hawaii
Herbivory/grazing/browsing
 
Urera kaalae
Hawaii
Ecosystem change / habitat alteration
Pest and disease transmission
 
Urocyon littoralis catalinae (Santa Catalina Island fox)
California
Ecosystem change / habitat alteration
 
Verbesina dissita (big-leaved crownbeard)
California
Herbivory/grazing/browsing
 
Vetericaris chaceorum (anchialine pool shrimp)
Hawaii
Ecosystem change / habitat alteration
 
Vicia menziesii (Hawaiian vetch)
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Vigna o-wahuensis (Oahu cowpea)
Hawaii
Herbivory/grazing/browsing
 
Viola chamissoniana subsp. chamissoniana (pamakani)
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Wilkesia hobdyi (dwarf iliau)
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 
Zanthoxylum dipetalum var. tomentosum
Hawaii
Ecosystem change / habitat alteration
Herbivory/grazing/browsing
 

Risk and Impact Factors

Invasiveness

Proved invasive outside its native range
Highly adaptable to different environments
Capable of securing and ingesting a wide range of food

Impact outcomes

Ecosystem change/ habitat alteration
Negatively impacts agriculture
Negatively impacts forestry
Negatively impacts animal health
Negatively impacts livelihoods
Reduced native biodiversity
Threat to/ loss of native species

Impact mechanisms

Competition - monopolizing resources
Competition (unspecified)
Pest and disease transmission
Herbivory/grazing/browsing
Interaction with other invasive species
Pathogenic

Uses

The ability of goat herds to survive in harsh environments has led to their introduction onto many islands, including Saint Helena Island, the Juan Fernandez Islands and Hawaii. They provided food for colonising people (specifically European colonisers and ship crews) (Campbell and Donlan 2005). Fishermen may have spread goats onto new islands, such as San Benitos (Mexico) and Pinta Island and Marchena Island (Galapagos Islands, Ecuador) (Campbell and Donlan 2005).
Goats were domesticated 10,000 years ago in the highlands of western Iran (Zeder and Hesse 2000, in Campbell and Donlan 2005). Goats are used for their fur and meat as well as for milk and cheese production. Goat meat is the most highly consumed meat source in the world. More goat’s milk is consumed than cow’s milk. Angora goats have long soft fur which is utilised to produce a soft silk-like fabric called mohair. In New Zealand, mohair prices are strong (2006) and farm gate returns are good. Although little is known of optimum farm management systems, it is considered that the farming of angora goats could contribute to the positive growth of the economy (Mohair NZ Business Plan 2006). The French are well known for utilising goat products for economic purposes; making cheese and other goat milk products (Canus Undated). Some cosmetic products have claimed to help eczema sufferers (Johnson 2006).

Uses List

Materials > Skins/leather/fur
Human food and beverage > Fresh meat
Human food and beverage > Milk and dairy products

Prevention and Control

Due to the variable regulations around (de)registration of pesticides, your national list of registered pesticides or relevant authority should be consulted to determine which products are legally allowed for use in your country when considering chemical control. Pesticides should always be used in a lawful manner, consistent with the product's label.
Management Information

Original text compiled by IUCN SSC Invasive Species Specialist Group (ISSG)

Goats have been eradicated from approximately 120 islands and there is hope that native communities will recover rapidly (Hamann 1979, 1993, in Campbell and Donlan 2005). The largest islands on which substantial goat populations have been removed are: Lanai Island (Hawaii), San Clemente (USA), Pinta Island and Santiago Island (Galapagos Islands, Ecuador) and Raoul Island (New Zealand) (Campbell and Donlan 2005; Cruz et al., 2009). In general, goat eradication management for islands larger than 500 hectares requires detailed planning and the use of specialised technology, equipment and personnel. Securing funds for eradication programmes may be an obstacle to goat control (Campbell and Donlan 2005). The recent eradication of feral goats from Santiago Island (58,465 ha) in <5 years cost US$6.1 million, and was achieved with a combination of techniques and technologies (Cruz et al. 2009).
The introduction of modern eradication technology has greatly improved the effectiveness of goat control programmes, making goat eradication more likely. Some control methods include the employment of aerial hunting (helicopter), specially trained goat-hunting dogs, Global Positioning Systems and Geographic Information System techniques as well as sterilised goats marked with radiotelemetry collars, called “Judas goats”, which gravitate to, and therefore detect, wild goat herds. Judas goats are used to find wild herds and are especially suitable for finding the last few survivors or to detect the presence of wild goats when it is uncertain whether they have been eradicated. Hunting dogs are particularly useful in situations when goat density is low and vegetation density is high. Aerial hunting is appropriate in situations where there is less ground cover and a higher density of goats. Eradication is always the better option when compared to short-term control. If short-term control is chosen, goats should be kept at low densities.
Although effective eradication technology is available, Benjamin et al. (2011) demonstrate the potential of strategic water management as a non-lethal technique to protect biodiversity from the impacts of feral goats. In the study area in New South Wales, feral goats were found to remain in close proximity to artificial water points. However, they were able to access and impact upon areas of conservation importance which were nearby to these water points. However, by using fencing, the distance the goats needed to travel from the water points to gain access to conservation areas was increased – this protected biodiversity without eradicating the goat population (although the population was locally reduced). This solution was favourable with local landholders some of whom harvest the feral goats.
Guidelines for managing the impact of feral goats have been developed under the Vertebrate Pest Program (VPP) Australia administered by the Bureau of Resource Sciences (BRS). The purpose of these guidelines is to assist in the development of cost-effective strategies to reduce the damage feral goats cause to production and conservation. Management techniques and strategies for feral goat management are recommended and illustrated by case studies. Deficiencies in knowledge, management and legislation are identified. Please follow this link to view and download Parkes, J., R Henzell & G Pickles, 1995. Managing Vertebrate Pests: Feral Goats

Reproduction

Oestrous cycle

Inter-oestrous interval is 20 days; oestrus duration is 40 hours. Ovulation occurs 30 hours after oestrus begins. Oestrus is seasonal only.

Links to Websites

NameURLComment
GISD/IASPMR: Invasive Alien Species Pathway Management Resource and DAISIE European Invasive Alien Species Gatewayhttps://doi.org/10.5061/dryad.m93f6Data source for updated system data added to species habitat list.
Global register of Introduced and Invasive species (GRIIS)http://griis.org/Data source for updated system data added to species habitat list.

Bibliography

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Atkinson, R., J. Renteria, W. Simbana. 2008. The Consequences of Herbivore Eradication on Santiago: Are We In Time to Prevent Ecosystem Degredation Again? In Galapagos Report 2007-8 FCD, PNG, and INGALA, Puerto Ayora, Galapagos, Ecuador.
Bellchambers, K., 2004. Improving the development of effective and humane trapping systems as a control method for feral goats in Australia.
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US Fish and Wildlife Service, 1998 . Kauai II: Addendum to the Recovery Plan for the Kauai Plant Cluster. In: Kauai II: Addendum to the Recovery Plan for the Kauai Plant Cluster.US Fish and Wildlife Service . 84 pp.
US Fish and Wildlife Service, 1998 . Recovery Plan for Oahu Plants. In: Recovery Plan for Oahu Plants.US Fish and Wildlife Service . 207 pp. + appendices.
US Fish and Wildlife Service, 1998 . Recovery Plan for the Nightingale Reed-warbler, Acrocephalus luscinia. In: Recovery Plan for the Nightingale Reed-warbler, Acrocephalus luscinia.US Fish and Wildlife Service . 62 pp.
US Fish and Wildlife Service, 1999 . Recovery Plan for Multi-Island Plants. In: Recovery Plan for Multi-Island Plants.US Fish and Wildlife Service . 206 pp. + appendices.
US Fish and Wildlife Service, 2002 . Endangered and Threatened Wildlife and Plants; Retention of Threatened Status for Argali in Kyrgyzstan, Mongolia, and Tajikistan. In: Federal Register, 67 ( 99 ) US Fish and Wildlife Service . 35942 - 35957 . http://www.gpo.gov/fdsys/pkg/FR-2002-05-22/pdf/02-12824.pdf#page=1
US Fish and Wildlife Service, 2003 . Endangered and Threatened Wildlife and Plants; Final Designation or Nondesignation of Critical Habitat for 95 Plant Species From the Islands of Kauai and Niihau, HI; Final Rule. In: Federal Register, 68 ( 39 ) US Fish and Wildlife Service . 9116 - 9164 . http://ecos.fws.gov/docs/federal_register/fr4053.pdf
US Fish and Wildlife Service, 2004 . Endangered and Threatened Wildlife and Plants; Listing the San Miguel Island Fox, Santa Rosa Island Fox, Santa Cruz Island Fox, and Santa Catalina Island Fox as Endangered. In: Endangered and Threatened Wildlife and Plants; Listing the San Miguel Island Fox, Santa Rosa Island Fox, Santa Cruz Island Fox, and Santa Catalina Island Fox as Endangered.US Fish and Wildlife Service . 19 pp.
US Fish and Wildlife Service, 2006 . Recovery Plan for the Kauai Cave Arthropods: The Kauai Cave Wolf Spider (Adelocosa anops) and the Kauai Cave Amphipod (Spelaeorchestia koloana). In: Recovery Plan for the Kauai Cave Arthropods: The Kauai Cave Wolf Spider (Adelocosa anops) and the Kauai Cave Amphipod (Spelaeorchestia koloana).US Fish and Wildlife Service . 64 pp.
US Fish and Wildlife Service, 2006 . Revised Recovery Plan for Hawaiian Forest Birds. In: Revised Recovery Plan for Hawaiian Forest Birds.US Fish and Wildlife Service . 622 pp.
US Fish and Wildlife Service, 2006 . Sibara filifolia (Santa Cruz Island Rockcress). 5-Year Review: Summary and Evaluation. In: Sibara filifolia (Santa Cruz Island Rockcress). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 24 pp.
US Fish and Wildlife Service, 2007 . Recovery Plan for Two Plants from Rota (Nesogenes rotensis and Osmoxylon mariannense). In: Recovery Plan for Two Plants from Rota (Nesogenes rotensis and Osmoxylon mariannense).US Fish and Wildlife Service . 86 pp.
US Fish and Wildlife Service, 2008 . 5-Year Review, Short Form Summary: Species Reviewed: Schiedea kauaiensis (no common name). In: 5-Year Review, Short Form Summary: Species Reviewed: Schiedea kauaiensis (no common name).US Fish and Wildlife Service . 6 pp.
US Fish and Wildlife Service, 2008 . 5-Year Review, Short Form Summary: Species Reviewed: Solanum incompletum (Popolo ku mai). In: 5-Year Review, Short Form Summary: Species Reviewed: Solanum incompletum (Popolo ku mai).US Fish and Wildlife Service . 5 pp.
US Fish and Wildlife Service, 2008 . Kulu`i (Nototrichium humile). 5-Year Review. In: Kulu`i (Nototrichium humile). 5-Year Review.US Fish and Wildlife Service . 11 pp.
US Fish and Wildlife Service, 2008 . Phyllostegia parviflora (no common name). 5-Year Review: Summary and Evaluation. In: Phyllostegia parviflora (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 12 pp.
US Fish and Wildlife Service, 2008 . Phyllostegia waimeae (no common name). 5-Year Review, Short Form. In: Phyllostegia waimeae (no common name). 5-Year Review, Short Form.US Fish and Wildlife Service . 6 pp.
US Fish and Wildlife Service, 2008 . Pritchardia kaalae (Lo'ulu). 5-Year Review: Summary and Evaluation. In: Pritchardia kaalae (Lo'ulu). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 11 pp.
US Fish and Wildlife Service, 2008 . Sanicula mariversa (no common name). 5-Year Review: Summary and Evaluation. In: Sanicula mariversa (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 10 pp.
US Fish and Wildlife Service, 2008 . Sierra Nevada Bighorn Sheep, Ovis canadensis californiana (= Ovis canadensis sierrae). 5-Year Review: Summary and Evaluation. In: Sierra Nevada Bighorn Sheep, Ovis canadensis californiana (= Ovis canadensis sierrae). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 40 pp.
US Fish and Wildlife Service, 2008 . Silene alexandri (no common name). 5-Year Review: Summary and Evaluation. In: Silene alexandri (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 10 pp.
US Fish and Wildlife Service, 2008 . Viola chamissoniana ssp. chamissoniana (Pamakani). 5-Year Review: Summary and Evaluation. In: Viola chamissoniana ssp. chamissoniana (Pamakani). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 11 pp.
US Fish and Wildlife Service, 2009 . 5-Year Review, Short Form Summary: Phyllostegia wawrana (no common name). In: 5-Year Review, Short Form Summary: Phyllostegia wawrana (no common name).US Fish and Wildlife Service . 5 pp.
US Fish and Wildlife Service, 2009 . 5-Year Review, Short Form Summary: Species Reviewed: Phyllostegia knudsenii (no common name). In: 5-Year Review, Short Form Summary: Species Reviewed: Phyllostegia knudsenii (no common name).US Fish and Wildlife Service . 6 pp.
US Fish and Wildlife Service, 2009 . 5-Year Review, Short Form Summary: Species Reviewed: Stenogyne campanulata (no common name). In: 5-Year Review, Short Form Summary: Species Reviewed: Stenogyne campanulata (no common name).US Fish and Wildlife Service . 6 pp.
US Fish and Wildlife Service, 2009 . 5-Year Review, Short Form Summary: Zanthoxylum dipetalum var. tomentosum (A`e). In: 5-Year Review, Short Form Summary: Zanthoxylum dipetalum var. tomentosum (A`e).US Fish and Wildlife Service . 7 pp.
US Fish and Wildlife Service, 2009 . Draft Revised Recovery Plan for the Mariana Fruit Bat or Fanihi (Pteropus mariannus mariannus). In: Draft Revised Recovery Plan for the Mariana Fruit Bat or Fanihi (Pteropus mariannus mariannus).US Fish and Wildlife Service . 98 pp. https://www.fws.gov/pacific/ecoservices/documents/100330.pdf
US Fish and Wildlife Service, 2009 . Phyllostegia mollis (no common name). 5-Year Review: Summary and Evaluation. In: Phyllostegia mollis (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 13 pp.
US Fish and Wildlife Service, 2009 . Remya mauiensis (Maui remya). 5-Year Review: Summary and Evaluation. In: Remya mauiensis (Maui remya). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 15 pp.
US Fish and Wildlife Service, 2009 . Schiedea nuttallii (no common name). 5-Year Review: Summary and Evaluation. In: Schiedea nuttallii (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 13 pp.
US Fish and Wildlife Service, 2009 . Solanum sandwicense (Popolo 'aiakeakua). 5-Year Review: Summary and Evaluation. In: Solanum sandwicense (Popolo 'aiakeakua). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 13 pp.
US Fish and Wildlife Service, 2009 . Tetramolopium lepidotum ssp. lepidotum (no common name). 5-Year Review: Summary and Evaluation. In: Tetramolopium lepidotum ssp. lepidotum (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 13 pp.
US Fish and Wildlife Service, 2009 . `O`u (Psittirostra psittacea). 5-Year Review: Summary and Evaluation. In: `O`u (Psittirostra psittacea). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 12 pp.
US Fish and Wildlife Service, 2010 . 5-Year Review Short Form Summary: Remya montgomeryi (no common name). In: 5-Year Review Short Form Summary: Remya montgomeryi (no common name) . US Fish and Wildlife Service . 7 pp. https://ecos.fws.gov/docs/five_year_review/doc3343.pdf
US Fish and Wildlife Service, 2010 . 5-Year Review, Short Form Summary: Species Reviewed: Poa mannii (Mann's bluegrass). In: 5-Year Review, Short Form Summary: Species Reviewed: Poa mannii (Mann's bluegrass).US Fish and Wildlife Service . 10 pp.
US Fish and Wildlife Service, 2010 . 5-Year Review, Short Form Summary: Species Reviewed: Pritchardia napaliensis (loulu palm). In: 5-Year Review, Short Form Summary: Species Reviewed: Pritchardia napaliensis (loulu palm).US Fish and Wildlife Service . 10 pp.
US Fish and Wildlife Service, 2010 . 5-Year Review, Short Form Summary: Species Reviewed: Pteralyxia kauaiensis (kaulu). In: 5-Year Review, Short Form Summary: Species Reviewed: Pteralyxia kauaiensis (kaulu).US Fish and Wildlife Service . 9 pp.
US Fish and Wildlife Service, 2010 . 5-Year Review, Short Form Summary: Species Reviewed: Remya kauaiensis (no common name). In: 5-Year Review, Short Form Summary: Species Reviewed: Remya kauaiensis (no common name).US Fish and Wildlife Service . 12 pp.
US Fish and Wildlife Service, 2010 . 5-Year Review, Short Form Summary: Species Reviewed: Schiedea helleri (no common name). In: 5-Year Review, Short Form Summary: Species Reviewed: Schiedea helleri (no common name).US Fish and Wildlife Service . 6 pp.
US Fish and Wildlife Service, 2010 . 5-Year Review, Short Form Summary: Species Reviewed: Schiedea kealiae (ma oli oli). In: 5-Year Review, Short Form Summary: Species Reviewed: Schiedea kealiae (ma oli oli).US Fish and Wildlife Service . 6 pp.
US Fish and Wildlife Service, 2010 . 5-Year Review, Short Form Summary: Species Reviewed: Schiedea membranacea (no common name). In: 5-Year Review, Short Form Summary: Species Reviewed: Schiedea membranacea (no common name).US Fish and Wildlife Service . 9 pp.
US Fish and Wildlife Service, 2010 . 5-Year Review, Short Form Summary: Species Reviewed: Schiedea stellarioides (no common name). In: 5-Year Review, Short Form Summary: Species Reviewed: Schiedea stellarioides (no common name).US Fish and Wildlife Service . 7 pp.
US Fish and Wildlife Service, 2010 . 5-Year Review, Short Form Summary: Wilkesia hobdyi (dwarf iliau). In: 5-Year Review, Short Form Summary: Wilkesia hobdyi (dwarf iliau).US Fish and Wildlife Service . 8 pp.
US Fish and Wildlife Service, 2010 . 5-year review, Short Form Summary: Zanthoxylum hawaiiense (a'e, manele). In: 5-year review, Short Form Summary: Zanthoxylum hawaiiense (a'e, manele).US Fish and Wildlife Service . 10 pp.
US Fish and Wildlife Service, 2010 . Determination of Endangered Status for 48 Species on Kauai and designation of Critical Habitat: Final Rule. In: Determination of Endangered Status for 48 Species on Kauai and designation of Critical Habitat: Final Rule.US Fish and Wildlife Service . i + 205 pp.
US Fish and Wildlife Service, 2010 . Higo Chumbo (Harrisia portoricensis). 5-Year Review: Summary and Evaluation. In: Higo Chumbo (Harrisia portoricensis). 5-Year Review: Summary and Evaluation . US Fish and Wildlife Service . 19 pp. http://ecos.fws.gov/docs/five_year_review/doc3078.pdf
US Fish and Wildlife Service, 2010 . Plantago princeps (laukahi kuahiwi). 5-Year Review: Summary and Evaluation. In: Plantago princeps (laukahi kuahiwi). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 19 pp.
US Fish and Wildlife Service, 2010 . Recovery Outline for the Kauai Ecosystem. In: Recovery Outline for the Kauai Ecosystem.US Fish and Wildlife Service . 38 pp. + 3 maps.
US Fish and Wildlife Service, 2010 . Schiedea apokremnos (maolioli). 5-Year Review: Summary and Evaluation. In: Schiedea apokremnos (maolioli). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 16 pp.
US Fish and Wildlife Service, 2010 . Sesbania tomentosa (ohai). 5-Year Review: Summary and Evaluation. In: Sesbania tomentosa (ohai). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 24 pp.
US Fish and Wildlife Service, 2010 . Silene lanceolata (no common name). 5-Year Review: Summary and Evaluation. In: Silene lanceolata (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 15 pp.
US Fish and Wildlife Service, 2010 . Spermolepis hawaiiensis (no common name). 5-Year Review: Summary and Evaluation. In: Spermolepis hawaiiensis (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 19 pp.
US Fish and Wildlife Service, 2010 . Tetramolopium filiforme (no common name). 5-Year Review: Summary and Evaluation. In: Tetramolopium filiforme (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 11 pp.
US Fish and Wildlife Service, 2010 . Verbesina dissita (Big-leaved crownbeard). 5-Year Review: Summary and Evaluation. In: Verbesina dissita (Big-leaved crownbeard). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 38 pp. https://ecos.fws.gov/docs/five_year_review/doc3559.pdf
US Fish and Wildlife Service, 2011 . 5-Year Review, Short Form Summary: Species Reviewed: Pritchardia aylmer-robinsonii (wahane). In: 5-Year Review, Short Form Summary: Species Reviewed: Pritchardia aylmer-robinsonii (wahane).US Fish and Wildlife Service . 9 pp.
US Fish and Wildlife Service, 2011 . 5-Year Review, Short Form Summary: Species Reviewed: Pritchardia munroi (lo'ulu). In: 5-Year Review, Short Form Summary: Species Reviewed: Pritchardia munroi (lo'ulu).US Fish and Wildlife Service . 11 pp.
US Fish and Wildlife Service, 2011 . 5-Year Review, Short Form Summary: Vigna o-wahuensis (no common name). In: 5-Year Review, Short Form Summary: Vigna o-wahuensis (no common name).US Fish and Wildlife Service . 10 pp.
US Fish and Wildlife Service, 2011 . Endangered and Threatened Wildlife and Plants; Listing 23 Species on Oahu as Endangered and Designating Critical Habitat for 124 Species. In: Federal Register, 76 ( 148 ) US Fish and Wildlife Service . 46362 - 46593 . http://www.gpo.gov/fdsys/pkg/FR-2011-08-02/pdf/2011-17162.pdf
US Fish and Wildlife Service, 2011 . Hawaiian Dark-rumped Petrel (Pterodroma phaeopygia sandwichensis). 5-Year Review: Summary and Evaluation. In: Hawaiian Dark-rumped Petrel (Pterodroma phaeopygia sandwichensis). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 16 pp.
US Fish and Wildlife Service, 2011 . Isodendrion longifolium (aupaka). 5-Year Review: Summary and Evaluation. In: Isodendrion longifolium (aupaka). 5-Year Review: Summary and Evaluation . US Fish and Wildlife Service . 28 pp. http://ecos.fws.gov/docs/five_year_review/doc3809.pdf
US Fish and Wildlife Service, 2011 . Panicum fauriei var. carteri (no common name). 5-Year Review: Summary and Evaluation. In: Panicum fauriei var. carteri (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 17 pp.
US Fish and Wildlife Service, 2011 . Phyllostegia mannii (no common name). 5-Year Review: Summary and Evaluation. In: Phyllostegia mannii (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 14 pp.
US Fish and Wildlife Service, 2011 . Schiedea haleakalensis (no common name). 5-Year Review: Summary and Evaluation. In: Schiedea haleakalensis (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 17 pp.
US Fish and Wildlife Service, 2011 . Schiedea hookeri (no common name). 5-Year Review: Summary and Evaluation. In: Schiedea hookeri (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 20 pp.
US Fish and Wildlife Service, 2011 . Schiedea lydgatei (no common name). 5-Year Review: Summary and Evaluation. In: Schiedea lydgatei (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 17 pp.
US Fish and Wildlife Service, 2011 . Schiedea sarmentosa (no common name). 5-Year Review: Summary and Evaluation. In: Schiedea sarmentosa (no common name). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 14 pp.
US Fish and Wildlife Service, 2011 . Urera kaalae (opuhe). 5-Year Review: Summary and Evaluation. In: Urera kaalae (opuhe). 5-Year Review: Summary and Evaluation.US Fish and Wildlife Service . 19 pp.
US Fish and Wildlife Service, 2012 . Endangered and Threatened Wildlife and Plants; Endangered Status for 23 Species on Oahu and Designation of Critical Habitat for 124 Species; Final Rule. In: Federal Register, 77 ( 181 ) US Fish and Wildlife Service . 57648 - 57862 . https://www.gpo.gov/fdsys/pkg/FR-2012-09-18/pdf/2012-19561.pdf
US Fish and Wildlife Service, 2012 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Synthliboramphus hypoleucus. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Synthliboramphus hypoleucus . US Fish and Wildlife Service . 24 pp. http://ecos.fws.gov/docs/candidate/assessments/2012/r8/B098_V01.pdf
US Fish and Wildlife Service, 2013 . Endangered and Threatened Wildlife and Plants; Determination of Endangered Species Status for 15 Species on Hawaii Island; Final Rule. In: Federal Register, 78 ( 209 ) US Fish and Wildlife Service . 64638 - 64690 . https://www.gpo.gov/fdsys/pkg/FR-2013-10-29/pdf/2013-24103.pdf
US Fish and Wildlife Service, 2013 . Endangered and Threatened Wildlife and Plants; Determination of Endangered Status for 38 Species on Molokai, Lanai, and Maui; Final Rule. In: Federal Register, 78 ( 102 ) US Fish and Wildlife Service . 32014 - 32065 . https://www.gpo.gov/fdsys/pkg/FR-2013-05-28/pdf/2013-12105.pdf
US Fish and Wildlife Service, 2013 . Endangered and Threatened Wildlife and Plants; Review of Native Species That are Candidates for Listing as Endangered or Threatened; Annual Notice of Findings on Resubmitted Petitions; Annual Description of Progress on Listing Actions. In: Federal Register, 78 ( 226 ) US Fish and Wildlife Service . 70104 - 70162 . https://www.gpo.gov/fdsys/pkg/FR-2013-11-22/pdf/2013-27391.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Gardenia remyi. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Gardenia remyi . US Fish and Wildlife Service . 17 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/Q33N_P01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Huperzia (= Phlegmariurus) stemmermanniae. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Huperzia (= Phlegmariurus) stemmermanniae . US Fish and Wildlife Service . 14 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/S02B_P01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus assimulans. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus assimulans . US Fish and Wildlife Service . 33 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/I0GQ_I01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus facilis. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus facilis . US Fish and Wildlife Service . 32 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/I0GY_I01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus hilaris. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus hilaris . US Fish and Wildlife Service . 31 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/I0HT_I01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus kuakea. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus kuakea . US Fish and Wildlife Service . 29 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/I0VM_I01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus longiceps. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus longiceps . US Fish and Wildlife Service . 33 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/I0HC_I01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus mana. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Hylaeus mana . US Fish and Wildlife Service . 30 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/I0VL_I01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Joinvillea ascendens ascendens. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Joinvillea ascendens ascendens . US Fish and Wildlife Service . 22 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/Q1XN_P01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Ochrosia haleakalae. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Ochrosia haleakalae . US Fish and Wildlife Service . 17 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/Q3ES_P01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Pseudognaphalium sandwicensium var. molokaiense. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Pseudognaphalium sandwicensium var. molokaiense . US Fish and Wildlife Service . 13 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/Q1WN_P01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Ranunculus hawaiensis. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Ranunculus hawaiensis . US Fish and Wildlife Service . 16 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/Q38H_P01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Schiedea pubescens. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Schiedea pubescens . US Fish and Wildlife Service . 14 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r1/Q2VZ_P01.pdf
US Fish and Wildlife Service, 2014 . U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Streptanthus bracteatus. In: U.S. Fish and Wildlife Service species assessment and listing priority assignment form: Streptanthus bracteatus . US Fish and Wildlife Service . 24 pp. http://ecos.fws.gov/docs/candidate/assessments/2014/r2/Q1R7_P01.pdf
US Fish and Wildlife Service, 2015 . Endangered and Threatened Wildlife and Plants; Endangered Status for 16 Species and Threatened Status for 7 Species in Micronesia. In: Federal Register, 80 ( 190 ) US Fish and Wildlife Service . 59424 - 59497 . https://www.gpo.gov/fdsys/pkg/FR-2015-10-01/pdf/2015-24443.pdf
US Fish and Wildlife Service, 2015 . Endangered and Threatened Wildlife and Plants; Endangered Status for 49 Species From the Hawaiian Islands. In: Federal Register, 80 ( 189 ) US Fish and Wildlife Service . 58820 - 58909 . https://www.gpo.gov/fdsys/pkg/FR-2015-09-30/pdf/2015-24305.pdf

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